Gebruiker:Joopwiki/Kladblok/Beenderknagers

Cm=Cretomorgus, Mt=Madagatrox, OA=Omorgus (Afromorgus), OH=Omorgus (Haroldomorgus), OO=Omorgus (Omorgus), Pn=Polynoncus, TP=Trox (Phoberus), TT=Trox (Trox). HT=holotype, AT=allotype, PT(s)=paratype(s), LT=lectotype, STs=syntypes. m=male, f=female. Au.=Australia, Ter.=Territory, RSA=Republic of South Africa. Invalid names are offset by emdashes. Checklist

Afromorgus Scholtz, 1986a: 362; type sp. Trox squalidus Olivier, 1789, original designation. Cretomorgus Nikolaev, 2007: 106, 215; type sp. Cretomorgus ikhbogdensis Nikolaev, 2007, original designation. Fossil (Early Cretaceous). Haroldomorgus Scholtz, 1986a: 362; type sp. Trox batesi Harold, 1872, monotypy. Madagatrox Pittino, 210: 75; type sp. Madagatrox ranotsaraensis Pittino, 2010: 77, monotypy. Omorgus Erichson, 1847: 111; type sp. Trox suberosus Fabricius, 1775, subsequent designation by Lacordaire (1856). Syn. Chesas Burmeister, 1876: 264; type sp. Trox pastillarius Blanchard, 1846, original designation. Syn. by Vaurie (1962: 109, 115). Syn. Lagopelas Burmeister, 1876: 265; type sp. Trox ciliatus Blanchard, 1846, original designation. Syn. by Vaurie (1962: 109, 115). Syn. Megalotrox Preudhomme de Borre, 1886: 59; type sp. Trox gigas Harold, 1872, subsequent designation by Scholtz (1982). Syn. by Vaurie (1962: 109, 115). Phoberus MacLeay, 1819: 137-138; type sp. Trox horridus Fabricius, 1775, monotypy. Polynoncus Burmeister, 1876: 264; type sp. Trox pedestris Harold, 1872, subsequent designation by Scholtz (1982). Trox Fabricius, 1775: 31; type sp. Scarabaeus sabulosus Linné, 1758, subsequent designation by Latreille (1810). Syn. Pseudotrox Robinson, 1948: 1; type sp. Trox laticollis LeConte, 1854, monotypy. Syn. by Vaurie (1955: 18). Species-group taxa acanthinus Harold, 1872: 154; TT; Mexico (Veracruz); STs – BMNH. —aciculeatus Robinson, 1940: 156; HTm – ANSP; = foveicollis (syn. by Vaurie 1955: 39). acinus (Scholtz, 1980a: 39); OA; ne. Tanzania; HTf – SMTD, PTf – UPSA. aculeatus Harold, 1872: 37; TP; “Caffraria”; LTm – MNHN (des. by Scholtz 1980a: 64). aeger (Guérin, 1844: 85); Pn; Argentina, Brazil, Paraguay, Uruguay (not in Chile, see Diéguez 2008); type – MNHN?; syn. leprosus. aequalis Say, 1831: 5; TT; Canada (Manitoba, New Brunswick, Nova Scotia, Ontario, Quebec), USA (e. states s. to South Carolina and w. to Texas and North Dakota; also Arizona, Colorado), Mexico (Durango, Chihuahua, Coahuila, Morelos, Nuevo León); Type(s) destroyed or MCZH? affinis Robinson, 1940: 158, as ssp. of aequalis; TT; USA (ne. states, also California, Iowa, Texas, possibly Kansas and Louisiana); HTm – ANSP. —aino Nakane and Tsukamoto, 1955: 92; HT – NSMT; = setifer CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 7 alatus (MacLeay, 1888: 907); OO; Australia (Northern Ter., Western Au.); HT – ANIC. alius Scholtz, 1986b: 50; OO; Australia (Western Au.); HT – QMBA. alpigenus Zhang, 1988: 234; TT; China (Xizang); HT – CASB. alternans (MacLeay, 1826: 3, 1827: 439); OO; Australia (Capital Ter., South Au., Victoria, Western Au.); HT – ANIC; syn. australasiae Germar, litigiosus. [Described as new twice, in 1826 and in a larger version of the work issued in 1827.] —alternans LeConte, 1854: 211 (n. praeocc., see lecontei); HT [=LT] – MCZH; = sonorae (syn. by Horn 1874: 8). —alternatus (Say, 1835: 179); = suberosus (syn. by Harold 1872: 120). amictus (Haaf, 1954b: 714); OO; Australia (South Au., Western Au.); HT – ISNB, PT – BMNH. amitinus (Kolbe, 1904: 294); OA; se. Kenya; LT – MNHB (des. by Scholtz 1980a: 38). —angulatus Fåhraeus, 1857: 381; LTm – NHRS (des. by Scholtz 1980a: 90); = squamiger (syn. by Haaf 1953: 326). —annexus (Kolbe, 1904: 293); type lost; = denticulatus (syn. by Haaf 1954a: 354). antiquus Wickham, 1909: 129; fossil, gen. inc. sedis; USA (Colorado: Florissant), late Eocene; HT – USNM (Dept. Invertebrate Paleontology). aphanocephalus Scholtz, 1986b: 18; OO; Australia (Queensland); HT – MVMA. aproximans Escalera, 1914: 158, as var. of confluens; TT; Morocco; LTm – IEEM (des. by Pittino 1983: 276); made available as ssp. of confluens by Balthasar (1936: 452). arcuatus Haaf, 1953: 323; TP; RSA (e. Cape); HTm – NHMB, PTs – BMNH, MNHN. —arenarius Fabricius, 1787: 18; type – ZMUC; = scaber (syn. by Harold 1872: 183). —arenarius Paykull, 1798: 80; = hispidus (syn. by Harold 1872: 165). —arenosus Gmelin, 1790: 1586; type lost; = scaber (syn. by Harold 1872: 183). argentinus (Harold, 1872: 143); Pn; Argentina (Córdova, Neuquén, Buenos Aires, Bahia Blanca); HTm – MNHN; syn. with gemmifer by Vaurie (1962: 120), reinstated by Pittino (1987: 384). aricensis (Gutiérrez, 1950: 63); Pn; Argentina, Bolivia, Chile, Peru; HTm – UCCH, PTs – BMNH. —asiaticus Faldermann, 1835a: 261; = niger (syn. by Pittino 2006: 80). —asmaraensis Balthasar, 1939: 32; HT – NMPC; = nigrociliatus (syn. by Haaf 1953: 328). asper LeConte, 1854: 215; OO; USA (from Rhode Island to Florida and w. to Nebraska, Kansas and Texas; also in Utah), Mexico (n. Sonora); HT – MCZH. —asperatus (MacLeay, 1888: 908); = crotchi (syn. by Blackburn 1904: 284). —asperrimus (MacLeay, 1888: 909); = semicostatus (syn. by Blackburn 1904: 284). asperulatus (Harold, 1872: 75); OA; Botswana, Namibia, RSA; LTm – MNHN (des. by Scholtz 1979a: 189). atrox LeConte, 1854: 214; TT; Canada (Alberta, British Columbia, Ontario, Saskatchewan), USA (w. states e. to Mississippi River; also Illinois, Indiana, Iowa, Michigan), Mexico (Durango); HT – MCZH. augustae (Blackburn, 1892: 39); OO; Australia (New South Wales, Northern Ter., Queensland, South Au., Victoria); HT – BMNH. australasiae (Erichson, 1842: 158); OO; Australia (all states). —australasiae (Germar, 1848: 189) [n. n. litigiosus Harold]; LT – MNHN; = alternans (syn. and LT des. by Scholtz 1986b: 46). baccatus (Gerstaecker, 1866: 49); OA; Kenya, Tanzania; LTm – MNHB (des. by Scholtz 1980a: 59). badeni (Harold, 1872: 83); OO; ne. Brazil, n. Colombia; LT – ISNB (des. by Scholtz 1990b: 1417). —barbarus Harold, 1872: 169; LTm – MNHN ; = granulipennis (syn. and LT des. by Pittino 2006: 80). —barbosus Laicharting, 1781: 31; = scaber (syn. by Harold 1872: 183). batesi (Harold, 1872: 126); OH; n. Argentina, Brazil, Paraguay; LT – ISNB (des. by Scholtz 1990b: 1418). —berytensis Petrovitz, 1980: 614; HTm – MHNG; = mixtus (syn. by Pittino 1983: 271). bifurcatus (Vaurie, 1962: 129); Pn; Argentina, Bolivia, Brazil, Paraguay; HTm + PTs – MNHB, PTs at other institutions; redescribed by Pittino (1987: 393). birmanicus (Arrow, 1927: 467); OO; Myanmar; HT – BMNH. borgognoi (Marchand, 1902: 332); OA; Chad, Mali, Mauritania; type lost. 8 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK borrei (Harold, 1872: 84); OO; Argentina, s. Bolivia, Brazil(?), Uruguay; HT + PT – ISNB. boucomonti Paulian, 1933: 205; TT; China (Fujian, Jiangxi, Yunnan); HT – MNHN. braacki Scholtz, 1980a: 87; TP; RSA (s. Cape); HTm + PTs – TMSA, PT – BMNH. brahminus Pittino, 1985: 338; TT; China (Xizang, Yunnan), India (Himachal Pradesh, Punjab, Sikkim, West Bengal), Malaya, n. Vietnam; HTm – BMNH, PTm – RPMI. brasiliensis (Vaurie, 1962: 134); Pn; Brazil, Paraguay; HTm – MZSP, PTs – AMNH and other institutions. brevicollis (Eschscholtz, 1822: 12); Pn; Argentina, Brazil, Chile, Colombia, Paraguay, Peru; ST(1) – BMNH; syn. lachrymosus. brincki Haaf, 1958a: 475; TP; Lesotho (Drakensberg Mtns.); HTm + PTs – MZLU, PTs – NHMB. brucki (Harold, 1872: 93); OO; Australia (Capital Ter., New South Wales, Queensland, Victoria); type – MNHN? bullatus (Curtis, 1845: 444); Pn; Argentina, Bolivia, Chile, Peru, Uruguay; HT – BMNH. burmeisteri Pittino, 1987: 380; Pn; Argentina (Mendoza); HTm + PTs – MNHN and other institutions. cadaverinus cadaverinus Illiger, 1802b: 44; TT; c. Europe, w. Russia, Siberia, Mongolia, Kyrgyzstan, Syria. cadaverinus komareki Balthasar, 1931: 130, as sp.; China (Heilongjiang), Mongolia, Siberia (Ussuri region); HT – NMPC. caffer caffer Harold, 1872: 41; TP; RSA (Cape, Free State); HT + PT – MNHB; Scholtz (1980a: 95) was unable to examine the MNHB types listed by Haaf (1953: 334) and designated a LT (MNHN), which is invalid. caffer lilianae Scholtz, 1980a: 96; Malawi, e. RSA, e. Zimbabwe; HTm + PTs – NMBZ, PTs – BMNH + TMSA + UPSA. cambeforti Pittino, 1985: 334; TT; China (Sichuan); HT + AT + PTs – MNHN. cambodjanus Pittino, 1985: 331; TT; Cambodia; HTf – MNHN; male described by Pittino and Kawai (2006). —canaliculatus Say, 1824: 278; type? – MCZH; = tuberculatus DeGeer (syn. by Harold 1872: 155). candezei (Harold, 1872: 113); OO; Argentina; type – MNHN?, ISNB? candidus (Harold, 1872: 98); OO; Australia (Capital Ter., New South Wales, Queensland, South Au., Victoria); LT – MNHN (des. by Scholtz 1986b: 28). capensis Scholtz, 1979b: 174; TT; RSA (s. Cape); HTm – TMSA, PTs – BMNH and other institutions. capillaceus Scholtz, 1990b: 1413; OO; Colombia (Bogotá); HTf – CMNH; male unknown. capillaris Say, 1823: 238; TT; Canada (Ontario), USA (ne. states to Tennessee, Missouri, Kansas; also Illinois, Indiana, Iowa, Texas, Utah); type? – MCZH. carinatus (Loomie, 1922: 135); OO; USA (s. Arizona, s. New Mexico, sw. Texas), Mexico (Chihuahua); HTm – USNM. carinicollis Scholtz, 1986b: 52; OO; Australia (Western Au.); HT – WAMP. —castelnaui (Lansberge, 1887: 201); = gigas (syn. by Blackburn 1904: 285). —chevrolati Harold, 1868: 86; = perlatus (syn. by Harold 1872: 161). chilensis (Harold, 1872: 137); Pn; w. Argentina, Chile; HT – MNHN. chinensis (Boheman, 1858: 52); OA; e. China, Taiwan, Japan, se. Asia, Indonesia (Java, Sumatra); HTm – NHRS; syn. obscurus. ciliatus (Blanchard, 1846: 190); OO; Argentina, Bolivia; type – MNHN. clathratus Reiche, 1861: 205; TT; France (Corsica), Italy (Sardinia). —concinnus Erichson, 1848: 932; = eversmanni. confluens Wollaston, 1864: 193; TT; Algeria, Canary Is., Libya, Morocco, Tunisia; HT – BMNH. —confluentus (Robinson, 1941a: 134); HTm – ANSP; = umbonatus (syn. by Vaurie 1955: 76). —conjunctus Petrovitz, 1975: 2; = sonorae (syn. by Pittino 2006: 28, 81). consanguineus (Péringuey, 1901: 466); OA; Namibia, Zaire, n. Zimbabwe; LT – SAMC (des. by Scholtz 1980a: 34). consimilis Haaf, 1953: 324; TP; Namibia, RSA; HT + AT – NHMB, PTs – BMNH. contractus Robinson, 1940: 153; TT; USA (sc. Texas); HTm – ANSP. —coracinus Gmelin, 1790: 1587; inc. sed.; patria? CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 9 costatus (Wiedemann, 1823: 30); OO; Australia (all states), se. Asia, s. China, Papua New Guinea, Solomon Is.; syn. montalbanensis, regularis, velutinus. cotodognanensis Compte, 1986: 69; TT; Spain. —crenatus (Olivier, 1789: no. 4: 7); = suberosus (syn. by Harold 1872: 120). cretaceus Nikolaev, 2007: 102f, 105, 206, 215; Trox inc. sed.; fossil (Early Cretaceous), Mongolia. cribrum Gené, 1836: 186; TT; France (Corsica), Italy (Sardinia). —cricetulus Ádám, 1994c: 37; = hispidus (syn. by Pittino 2006: 28). crotchi (Harold, 1871: 116); OO; Australia (New South Wales, Northern Ter., Queensland, South Au., Victoria, Western Au.); LT – MNHN (des. by Scholtz 1986b: 26); syn. asperatus, eremita. curvipes (Harold, 1872: 90); OO; Australia (Northern Ter., Queensland, Western Au.); HT – MNHN. cyrtus Haaf, 1953: 333; TP; Lesotho, RSA; HTm + PTs – MNHN; syn. fumarius. —damarinus (Péringuey, 1901: 469); LT – SAMC; = rusticus (syn. and LT des. by Scholtz 1979a: 191). —demaisoni Reitter, 1904: 156; LT – HNHM; = transversus (syn. by Pittino 2006: 28). demarzi (Haaf, 1958: 1081); OO; Australia (Northern Ter., Queensland), w. Irian; HT – NHMB. dentibius Robinson, 1940, see ssp. of spinulosus. denticulatus (Olivier, 1789: 13); OA; Djibouti, Ethiopia, Kenya, Somalia, Sudan, Tanzania, Uganda; type lost; syn. annexus, neumanni, ?verrucosus (see separate entry). —denticulatus (Beauvois, 1805: 176); = suberosus (syn. by Harold 1872: 120). —denticulatus (Blanchard, 1846: 189), praeocc. by denticulatus Olivier; = pedestris Harold, n. n. desertorum (Harold, 1872), see ssp. of niloticus. dhaulagiri Paulus, 1972a: 55; TT; Nepal; HTm – FISF. diffluens (Vaurie, 1962: 138); Pn; Chile (Santiago); HTf – CASC. dilaticollis (MacLeay, 1888: 907); OO; Australia (Northern Ter., Queensland, Western Au.); LT – ANIC (des. by Scholtz 1986b: 22). discedens (Haaf, 1954a: 359); OA; e. Kenya, se. Somalia, Tanzania; HTm – NHMV. dohrni (Harold, 1871: 112); OO; Australia (Western Au.); LT – MNHN (des. by Scholtz 1986b: 10). doiinthanonensis Masumoto, 1996: 9; TT; nw. Thailand; HTm – OWUT. ecuadorensis (Vaurie, 1962: 133); Pn; Ecuador, Peru ; HTm + PTs – ZMUC, and PTs at other institutions. elderi (Blackburn, 1892: 37); OO; Australia (South Au.: Ooldea); LTf – BMNH (des. by Scholtz 1986b: 12); male unknown. elevatus (Harold, 1872: 73); OA; s. Angola, Namibia; LTm – MNHN (des. by Scholtz 1979a: 190); syn. gravidus. elongatus (Haaf, 1954b: 706); OO; “Northern Australia”; HTm – MNHN; known only from HT. endroedyi (Scholtz, 1979a: 185); OA; s. Angola, nw. Namibia; HTm + PTs – TMSA. —eremita (Blackburn, 1892: 38); LT – BMNH; = crotchi (syn. and LT des. by Scholtz 1986b: 26). —erinaceus LeConte, 1854: 212; LT – MCZH; = variolatus (syn. and LT des. by Robinson 1940: 151). —erlangeri (Kolbe, 1904: 293); LTm – MNHB (des. by Scholtz 1980a: 55); = niloticus niloticus (syn. by Haaf 1954a: 345). erugatus Scholtz, 1990b: 1425; Pn; Argentina; HTf – FMLT; male unknown. euclensis (Blackburn, 1892: 39); OO; Australia (Capital Ter., New South Wales, Queensland, South Au., Victoria, Western Au.); LT – BMNH (des. by Scholtz 1986b: 44). eversmanni Krynicki, 1832: 123; TT; Europe, w. Russia, Siberia, Kazakhstan, Kyrgyzstan, Turkmenistan, Uzbekistan, Iran, China (Xinjiang). eximius Faldermann, 1835b: 368; TT; China (Nei Mongol), Mongolia. expansus (Arrow, 1900: 22); OA; ec. Somalia; HTm – BMNH; known only from HT. eyrensis (Blackburn, 1904: 293); OO; Australia (South Au., Western Au.); LT – BMNH (des. by Scholtz 1986b: 47). fabricii Reiche, 1853: 89; TT; Italy (Sicily), Portugal, Spain, Malta, Algeria, Libya, Morocco, Tunisia, Egypt. fascicularis fascicularis Wiedemann, 1821: 129; TT; RSA (s. Cape, n. Transvaal mtns.), e. Zimbabwe; LT – MNHB (des. by Scholtz 1980a: 78). 10 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK fascicularis rowei Scholtz, 1980a: 78; RSA (Free State, Natal mtns.); HTm + PTs – TMSA, PTs – BMNH. fascifer LeConte, 1854: 213; TT; Canada (British Columbia), USA (Pacific nw. from San Francisco to Canadian border); HT – MCZH. —femoratus (DeGeer, 1774: 269), as Scarabaeus; = sabulosus sabulosus (syn. by Harold 1872: 175). —fenestrellus (Balthasar, 1939: 34); type lost; = melancholicus (syn. by Scholtz 1979a: 188); revalidated by Pittino (2006: 27) without explanation. —fenestratus (Harold, 1872: 97); = subcarinatus (syn. by Haaf 1958b: 1079). floridanus Howden and Vaurie, 1957: 1; TT; USA (Florida); HTm + PTs – AMNH, ATf – CMNO, PTs at other institutions. formosanus Nomura, 1973: 37; TT; South Korea, Taiwan; HTm – NSMT. foveicollis Harold, 1872: 181; TT; Canada (Ontario), USA (e. states w. to Kansas, Oklahoma, Texas); type – MNHN?; syn. aciculeatus. foveolatus (Boheman, 1860: 114); OA; Namibia; LTm + PLTs – NHRS (des. by Scholtz 1979a: 184); syn. massalis; Paulian (1981: 7) reported this species from Madagascar, but his habitus photo (fig. 3) is not foveolatus and his drawing of the aedeagus (fig. 8d) appears to have been copied from Haaf (1954a: fig. 18). frater Pittino, 2005: 74; OA; India, Pakistan, Sri Lanka; HTm – RPMI, PTs – RPMI, SKTJ. freyi (Haaf, 1954a: 370); OA; Namibia, nw. RSA; HTm + PTs – NHMB, and PTs at other institutions. frontera Vaurie, 1955: 41; TT; USA (s. Texas); HTm + ATf + PTs – TAMU, and PTs at other institutions. fujiokai Ochi, 2000; see ssp. of sabulosus. —fumarius Haaf, 1953: 333; HTm – NHMB (syn. by Scholtz 1980a: 97). —furcifer Pittino, 1987: 394; HTm + PTs – HNHM, and PTs at other institutions; = bifurcatus (syn. by Scholtz 1990b: 1441). fuliginosus (Robinson, 1941a: 134); OO; USA (s. Texas), Mexico (Coahuila, Tamaulipas, Veracruz, Yucatán), Guatemala; HT – USNM. funestus (Lansberge, 1886: 94); OA; “Africa”; type lost; undocumented species, which the original description appears to place in the gemmatus or baccatus group of Omorgus (Afromorgus) – see Scholtz 1980a: 99, 1986a: 362. —furcillifer Pittino, 1987: 396; HTm – RPMI; = parafurcatus (syn. by Scholtz 1990b: 1441). galapagoensis (Van Dyke, 1953: 124); Pn; n. Galapagos (Darwin + Wolf Is.); HTf – CASC. gansuensis Ren, 2003: 110; TT; China (Gansu); HT – CASB? gemmatus (Olivier, 1789: 7); OA; Arabia, Chad, Djibouti, Egypt, Eritrea, Ethiopia, Gambia, Ghana, Guinea, Mali, Mauritania, Niger, Nigeria, Senegal, Somalia, Sudan; type lost. gemmifer (Blanchard, 1846: 187); Pn; Argentina, Brazil, Chile, Uruguay; HT – MNHN; syn. argentinus. —gemmiferus (Blanchard, 1846: 187); = gemmifer. —gemmiferus (Blanchard, 1846: 188), n. praeocc.; = guttifer. gemmingeri (Harold, 1872: 134); Pn; Argentina, Bolivia, Brazil, Paraguay, Uruguay (not in Chile, see Diéguez 2008); HT – MNHN. gemmulatus Horn, 1874: 8; TT; USA (Arizona, California, New Mexico, South Dakota), Mexico (n. Baja); LT – MCZH (des. by Robinson 1940: 151). genieri Scholtz, 1991b: 112; OA; Burkina Faso; HTm – LEMQ; known only from HT. gibberosus Scholtz, 1990b: 1428; Pn; Chile (Bíobio, Malleco); HTm – USNM, PTs – CMNO + UPSA. —gibbus (Olivier, 1789: no. 4: 13); type lost; = suberosus (syn. by Harold 1872: 120). gigas (Harold, 1872: 48); OO; Australia (South Au., Western Au.); type lost; syn. castelnaui. —glaber (Scholtz, 1980a: 36); OA; s. Tanzania; HTm – ZSBS; = lindemannae (syn. by Pittino 2005: 75). —globulatus (Fairmaire, 1883: 490); type – MNHN?; = hemisphaericus (syn. by Berg 1884: 98). —gonoderus (Fairmaire, 1901: 135); HT – MNHN; = squalidus (syn. by Paulian 1937: 134). gordoni (Steiner, 1981: 205); Pn; Peru (Cuzco); HT + AT + PTs – USNM. —graecus Mulsant and Rey, 1871: 223; = transversus. granulatus (Herbst, 1783: 12); OA; Afghanistan, India (Himachal Pradesh, Sikkim, West Bengal, c. + s. states), Pakistan; type lost. CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 11 —granulatus Fabricius, 1801: 110; HT – ZMUC; = fabricii (syn. by Pittino 2006: 80). granuliceps (Haaf, 1954b: 703); OO; “Australia”; HTf – MNHN; known only from HT. —gravidus (Péringuey, 1901: 470); LT – SAMC; = elevatus (syn. and LT des. by Scholtz 1979a: 190). granulipennis Fairmaire, 1852: 83; TT; Caucasus, Italy (Sicily), Spain (incl. Canary Is.), Gibraltar, Algeria, Egypt (incl. Sinai), Libya, Morocco, Tunisia, Iran, Israel, Lebanon, Syria; LT – MNHN (des. Pittino 1983: 268); syn. barbarus. gunki Scholtz, 1980a: 86; TP; RSA (s.-sw. Cape); HTm – TMSA, PTs – PPRI, UPSA. guttalis (Haaf, 1954a: 336); OA; Ghana, Mali, Nigeria, Senegal; HTm – NHMB. guttifer (Harold, 1868: 86), n. n. pro T. gemmiferus (Blanchard 1846: 188), praeocc. by T. gemmiferus (Blanchard 1846: 187); Pn; Argentina, Chile, Paraguay; HT – MNHN. haafi (Vaurie, 1962: 122); Pn; s. Argentina; HTm – MNHN, PTs – MNHB + MNHN + NHMB. haagi (Harold, 1872: 59); OA; India (West Bengal), Pakistan; HT – MNHN; regarded as a ssp. of niloticus by Haaf (1954a: 348) and Scholtz (1982: 12), elevated to species by Pittino (2006: 26, 27, 79). hamatus Robinson, 1940: 153; TT; Canada (Ontario), USA (e. states s. to Georgia + Tennessee and w. to Kansas + Nebraska; also Oklahoma, Texas); HTm – ANSP. —haroldi Flach, 1879: 155; type? – NMPC; = perrisii. hemisphaericus (Burmeister, 1876: 253); Pn; s. Argentina, s. Chile; ST – MACN?; syn. globulatus. hispanicus Harold, 1862a: 161; TT; France (Pyrenees), Portugal, Spain; HT – MNHN. hispidus (Pontoppidan, 1763: 431), as Scarabaeus; TT; Europe, Canary Is., Iran, Syria, Turkey; type lost; syn. arenarius Paykull (hom.), cricetulus, insularis Schneider (hom.), luridus Rossi (hom.), sabulosus Preyssler (hom.). —hispidus Paykull, 1798: 81; = scaber (syn. by Harold 1872: 183). horiguchii Ochi and Kawahara, 2002: 54; TT; Japan, North Korea. horridus Fabricius, 1775: 818; TP; RSA (sw. Cape); ST(2) – ZMUC; syn. pectinatus, silphoides. howdenorum Scholtz, 1986b: 14; OO; Australia (Western Au.); HT – SAMA, PT – BMNH. howelli (Howden and Vaurie, 1957: 4); OO; USA (ec. Florida, Texas); HTm – CMNO, PTm – INHS; presumed to occur through much of the range of its host bird Caracara (Polyborus cheriway), as far south as the Guianas and Peru.

• Cretomorgus ikhbogdensis Nikolaev, 2007: 107f, 215; Cm; fossil (Early Cretaceous), Mongolia.

inadai Ochi, Kawahara and Inagaki, 2008: 135; TT; Japan (Yakushima I.); HTm – OMNH [= opacotuberculatus?]. inclusus (Walker, 1858: 208); OA; China (Hainan), India, Sri Lanka; HTm – BMNH. —incultus (Fåhraeus, 1857: 377); LT – NHRS (des. by Scholtz 1979a: 187), PLT(1) – BMNH; = squalidus (syn. by Harold 1872: 108). indicus (Harold, 1872: 102); OA; India; “Typenmaterial” – MNHN. indigenus Scholtz, 1990b: 1411; OO; Galapagos (Española I.); HTm – CMNO, PTs – AMNH, BMNH and other institutions. ineptus Balthasar, 1931: 131; TT; e. Siberia, Mongolia; HT[ST?] – NMPC. inermis Pittino, 2005: 77; OA; s. India, Sri Lanka, s. Vietnam; HTm + PTs – RPMI. inflatus (Loomie, 1922: 135); OO; USA (Arizona, New Mexico, Texas), Mexico (Chihuahua); HTm – USNM. insignicollis (Blackburn, 1896: 265); OO; Australia (New South Wales, Northern Ter., Western Au.); HT – MVMA. insignis (Haaf, 1954a: 344); OA; s. Angola, Namibia; HTm – NHMB, PTs – TMSA. —instabilis Haaf, 1953: 331; HTm – NHMB, PT – BMNH; = sulcatus (syn. by Scholtz 1979a: 193). insularis Chevrolat, 1864: 416; TT; Cuba; HT – ANSP? [allegedly in Poey Coll., but not found by Robinson (1940)]. —insularis Schneider, 1898: 69; = hispidus. —integer LeConte, 1854: 216; LT – MCZH (des. by Robinson 1940: 151); = punctatus (syn. by Horn 1874: 5). iranicus Petrovitz, 1980: 613; TT; Iran; HTf – MHNG. italicus (Reiche, 1853: 89); OA; China (“Kwangu”), India (Himachal Pradesh, Kashmir, Sikkim, West Bengal), Pakistan, Italy (intro.); HT – MNHN. 12 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK jeanae Scholtz, Inward and Kerley, 2007: 98; TT; Thailand; HTm – BMNH; known only from HT. juglans (Ratcliffe, 1978: 299); Pn; Brazil, Guyana; HT + AT – INAM, PTs – AMNH and other institutions. kerleyi Masumoto, 1996: 10; TT; nw. Thailand; NSMT. kiuchii Masumoto, 1996: 11; TT; Laos, Thailand; HT – OWUT. klapperichi Pittino, 1983: 280; TT; Ukraine (Crimea), Greece, Cyprus, Turkey, Israel, Jordan, Syria, Iran, Iraq; HT + AT – MHNG, PTs – BMNH. komareki Balthasar, 1931, see ssp. of cadaverinus. koreanus Kim, 1991: 136; TT; South Korea. kyotensis Ochi and Kawahara, 2000: 53; TT; Japan. —lachrymosus (Curtis, 1845: 455); HT – BMNH; = brevicollis (syn. by Harold 1872: 145). lama Pittino, 1985: 329; TT; China (se. Xizang); HTm – BMNH. —larvatus (Gerstaecker, 1873: 309); LT – DEIM (des. by Scholtz 1979a: 188); = melancholicus (syn. by Haaf 1954a: 379). laticollis LeConte, 1854: 213; TT; USA (Arkansas, Indiana, Iowa, Massachusetts, Nebraska, New Jersey, New York, Pennsylvania); HT – MCZH. —lecontei Harold, 1872: 156, n. n. pro alternans LeConte, n. praeocc.; = sonorae (syn. by Horn 1874: 8). leonardii Pittino, 1983: 277; TT; Portugal, Spain, Algeria, Libya, Morocco, Tunisia, Egypt, Israel, Syria; HTm – MSNM, PTm – RPMI. —leprosus (Blanchard, 1846: 188); HT – MNHN; = aeger (syn. by Harold 1872: 132). levis Haaf, 1953: 325; TP; RSA (Cape, Free State); HTf – BMNH, PTs – ISNB. lilianae Scholtz, 1980, see ssp. of caffer. lindemannae (Petrovitz, 1975: 1); OA; Tanzania, e. RSA; HTm – ZSBS; syn. glaber. —litigiosus (Harold, 1872: 88) (n. n. pro australasiae Germar, n. praeocc.); = alternans (syn. by Scholtz 1986b: 46). litoralis Pittino, 1991: 77; TT; Albania, Bosnia-Hercegovina, Croatia, Montenegro, Serbia, Italy, Malta, Turkey; HT + AT – RPMI, PTs – BMNH, NMPC and other institutions. lobicollis (Arrow, 1927: 466); OA; Myanmar, Thailand; STs – BMNH. longitarsis (Harold, 1872: 136); Pn; Argentina, Chile; HT – MNHN. loxus (Vaurie, 1955: 58); OO; Mexico (Veracruz), Costa Rica, Argentina, Brazil, Colombia; HTf – MCZH; presumed to occur continuously from Mexico to Argentina. lucidus Pittino, 2010: 80; OO; Argentina (Buenos Aires); HTf – MSNM; known only from HT. lugubris (Haaf, 1954a: 338); OA; se. Kenya, Tanzania; HTm – NHMV. luridus Fabricius, 1781: 496; TP; Namibia, RSA (w. Cape); LT – ZMUC (des. by Scholtz 1979a: 194). —luridus Rossi, 1790: 17; = hispidus. lutosus (Marsham, 1802: 25), as Scarabaeus; TT, “England”; type lost, sp. inc. sed. —madagascariensis (Fairmaire, 1871: 34); HT – MNHN; = melancholicus (syn. by Harold 1872: 108). maindroni Pittino, 2005: 78; OA; Pakistan; HTm + ATf + PTs – MNHN, PT – RPMI. maissouri (Haaf, 1954a: 384), as ssp. of T. inclusus; “Vorder-Indien (Maissour, Shimoga), Surada und Burma”; HTm + PTs – MNHN; elevated to species by Pittino (2006: 27). mandli Balthasar, 1931: 128; TT; e. Siberia, China (Fujian, Heilongjiang), Japan, South Korea. HT – NMPC, syn. nishijimai. —manilensis (Schultze, 1915: 271); = suberosus (syn. by Haaf 1954b: 739). mariae Scholtz, 1986b: 13; OO; Australia (Western Au.); HT – WAMP. mariettae Scholtz, 1986b: 20; OO; Australia (Northern Ter.); HT – ANIC. marshalli (Haaf, 1957a: 694); OO; Australia (n. Queensland); HTf – NHMB; known only from HT redescribed by Pittino and Kawai (2010). martini Reitter, 1892: 153, as var. of granulipennis; TT; Spain, Algeria, Libya, Tunisia; LTf – DEIM (des. by Pittino 1983: 274); made available as ssp. of granulipennis by Balthasar (1936: 452). —massaicus (Kolbe, 1904: 293); LT – MNHB (des. by Scholtz 1980a: 55); = niloticus niloticus (syn. by Haaf 1954a: 345). —massalis (Harold, 1880: 156); type lost; = foveolatus (syn. by Haaf 1954a: 360). CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 13 matsudai Ochi and Hori, 1999: 37; TT; Japan (Amami-Ôshima). matsumurai Miyake and Yamaya, 1995, see ssp. of uenoi. maurus Herbst, 1790: 26; inc. sed.; patria? melancholicus (Fåhraeus, 1857: 378); OA; Angola, Botswana, Cameroon, CAR, Chad, Comoros, Eritrea, Ethiopia, Gambia, Ghana, Guinea, Ivory Coast, Kenya, Madagascar, Malawi, Mali, Mozambique, Namibia, Nigeria, RSA, Rwanda, Senegal, Sierra Leone, Sudan, Tanzania, Togo, Uganda, Zaire, Zambia, Zimbabwe; LT – NHRS (des. by Scholtz 1979a: 187); syn. fenestrellus, larvatus, madagascariensis [see note under fenestrellus]. mentitor (Blackburn, 1896: 266); OO; Australia (Northern Ter., South Au., Western Au.); HT – MVMA, “one type” – BMNH. mictlensis Deloya, 1995: 153; OO; Mexico (Puebla); HT + AT – IEXM. miliarius Gmelin, 1790: 1587; TP; RSA (Cape); type lost; undocumented species, which the original description appears to place in the sulcatus group of Trox (Phoberus) – see Scholtz 1980a: 99. mirabilis Pittino, 1987: 391; Pn; Argentina, Chile; HTm – RPMI. mitis Balthasar, 1933: 81; TT; e. Siberia, China (Beijing, Heilongjiang), Japan, South Korea; LT – NMPC (des. by Pittino 2006: 28). mixtus Harold, 1872: 167, as var. of granulipennis; TT; Cyprus, Turkey, Israel, Lebanon, Syria; LTm – MNHN (des. by Pittino 1983: 271); syn. berytensis Petrovitz; made available as sp. by Pittino 1983: 271 (as ssp. in 1985). mollis (Arrow, 1927: 468); OO; Indonesia (Borneo, Sumatra), Malaya; LT – BMNH (des. by Scholtz 1986b: 38) + PLT(1). monachus (Herbst, 1790: 25); OO; USA (e. states from the Great Lakes to Florida and w. to Nebraska, Kansas, Oklahoma and e. Texas); HT – MNHB; syn. pustulatus, tuberculatus Beauvois. —montalbanensis (Schultze, 1915: 272); = costatus (syn. by Haaf 1954a: 390). montanus Kolbe, 1891: 22; TP; Kenya, Tanzania, Uganda; LT – BMNH (des. by Scholtz 1980a: 77). monteithi Scholtz, 1986b: 58; OO; Australia (e. Queensland: Kroombit); HT – QMBA. —monticola Robinson, 1940: 158, as ssp. of variolatus; HT – ANSP; = variolatus (syn. by Vaurie 1955: 53). —morsus LeConte, 1854: 216; HT – MCZH; = punctatus (syn. by Horn 1874: 5). morticinii (Pallas, 1781: 11), as Scarabaeus; TT; sw. Russia, w. to e. Siberia, Kazakhstan, Kyrgyzstan, Turkmenistan, China (Xinjiang); syn. pinguis. mutabilis (Haaf, 1954a: 376); OA; Botswana, w. Ethiopia, Kenya, Malawi, Mozambique, e. RSA, Tanzania, Uganda, Zaire; HT – NHMB, PTs – BMNH. mutsuensis Nomura, 1937: 85; TT; Japan; HTm – NSMT. nama Kolbe, 1908: 125; TP; s. Namibia, RSA (nw. Cape); LTm – MNHB (des. by Scholtz 1979a: 195). nanningensis Pittino, 2005: 71; OO; China (Guangxi); HTm – RPMI; known only from HT. nanniscus Péringuey, 1901: 458; TT; RSA (e. Cape); LT – SAMC (des. by Scholtz 1979b: 177). nasutus Harold, 1872: 34; TT; RSA (sw. Cape); HTm – MNHN. natalensis natalensis Haaf, 1954c: 57; TT; RSA (se. Natal); HTm – DMSA. natalensis quadricostatus Scholtz, 1980a: 72; RSA (e. Transvaal); HTm – TMSA, PT – UPSA. natalensis schaborti Scholtz, 1980a: 72; RSA (e. Transvaal); HTm – TMSA. necopinus Scholtz, 1986c: 29, as TT; TP; nw. Zambia; HTm + PTs – BMNH; closest to T. squamiger in the sulcatus group, which Scholtz (1986a: 361) transferred to Trox (Phoberus). —neumanni (Kolbe, 1904: 293); LTm – MNHB (des. by Scholtz 1980a: 51); = denticulatus (syn. by Haaf 1954a: 354). neuquen (Vaurie, 1962: 123); Pn; Argentina, Chile; HTm + PTm – MNHB, PTf – CMNO. —nidicola Bonnaire, 1881: 63; = perrisii. niger Rossi, 1792: 9; TT; Europe, Siberia, Kazakhstan, Kyrgyzstan, Turkey; NTf – MNHB (des. by Pittino 1991: 65); syn. asiaticus. —nigrescens (Robinson, 1941a: 134); HTm – ANSP; = tesselatus (syn. by Vaurie 1955: 7). nigrociliatus nigrociliatus Kolbe, 1904; 292; TP; Ethiopia (highlands); LT – MNHB (des. by Scholtz 1980a: 92); syn. asmaraensis. nigrociliatus nyansanus Haaf, 1953: 330; Kenya, Rwanda, e. Zaire (lake areas); HTm – MNHB. nigrociliatus nyassicus Haaf, 1953: 330; e. Tanzania (montane); HTm – MNHB. 14 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK nigroscobinus Scholtz, 1986b: 20; OO; Australia (Western Au.); HTf – ANIC; male unknown. niloticus niloticus (Harold, 1872: 79); OA; Djibouti, Ethiopia, Kenya, Madagascar, Somalia, Sudan, Tanzania Uganda [S of 15ºN]; LTf – MNHN (des. by Scholtz 1980a: 55); syn. erlangeri, massaicus. niloticus desertorum (Harold, 1872: 77), as sp., demoted to ssp. by Haaf (1954a: 347); Arabia, Egypt, Eritrea [N of 15ºN]; LTf – MNHN (des. by Scholtz 1980a: 56); syn. philbyi. niponensis Lewis, 1895: 387; Japan, Taiwan, Nepal, + Oriental; TT; HT – BMNH; syn. with T. scaber by Arrow (1912: 61), reinstated by Pittino (2006: 80) without explanation. —nishijimai Nakane and Tsukamoto, 1955: 90; = mandli. —nobilis (Wollaston, 1867: 93); = suberosus (syn. by Harold 1872: 120). nocheles Scholtz, 1990b: 1412; OO; Argentina (Río Negro); HTm + PTm – LACM. nodicollis (MacLeay, 1888: 908); OO; Australia (Western Au.); LT – ANIC (des. by Scholtz 1986b: 56); doubtful species that may prove to be O. subcarinatus. nodosus (Robinson, 1940: 152); OO; USA (New Mexico, w. Texas), Mexico (n. Chihuahua); HTm – ANSP. nodulosus Harold, 1872: 165, as var. of hispidus; TT; France (Corsica), Italy (Sardinia); LTm – DEIM (des. by Pittino 1991: 69); made available as ssp. by Schmidt (1936: 76). nohirai Nakane, 1954: 10; TT; Japan; HT – NSMT. —nomadicus (Scholtz, 1980b: 138); = verrucosus (syn. by Pittino 2006: 28). —novaecaledoniae (Balthasar, 1966: 184); HT?, PTs(4) – NMPC; = suberosus (syn. by Scholtz 1986b: 54). nyansanus Haaf, 1953, see ssp. of nigrociliatus. nyassicus Haaf, 1953, see ssp. of nigrociliatus. obesus (Scholtz, 1980a: 30); OA; Djibouti, Ethiopia, Kenya, Tanzania, Uganda; HT + PTs – BMNH. —obscurus (Waterhouse, 1875: 98); HT – BMNH; = chinensis (syn. by Arrow 1927: 466). —occidentalis Pittino, 1987: 395, as ssp. of parafurcatus; = parafurcatus s. str. (syn. by Scholtz 1990b: 1441); HTm – MNHN, PTm – RPMI. —oligonus (Loomis, 1922: 133); HT – USNM; = texanus (syn. by Vaurie 1955: 75). omacanthus (Harold, 1872: 64); OA; India (Sikkim, Uttar Pradesh, West Bengal, c. states); type lost? opacotuberculatus Motschulsky, 1860: 41; TT; e. Siberia, China (Heilongjiang), Taiwan, Japan, Korean peninsula; type – ZMUM? —opacus Felsche, 1909: 450; LT – SMTD (des. by Scholtz 1980a: 74); = pusillus (syn. by Haaf 1953: 341). —oustaleti Scudder, 1879: 178B; n. dubium; fossil (Eocene), Nine-Mile Creek, BC, Canada. ovalis (Haaf, 1957b: 117); OO; Australia (Northern Ter.); HTf – MVMA; male unknown. —ovatus (Beauvois, 1805: 175); = suberosus (syn. by Harold 1872: 120). pampeanus (Burmeister, 1876: 255); Pn; Argentina; ST(4) – MACN. parafurcatus Pittino, 1987: 395; Pn; Argentina, Brazil; HT + PTs – MNHN; syn. furcillifer, parafurcatus occidentalis. parvicollis Scholtz, 1986b: 35; OO; Australia (New South Wales, Northern Ter., Victoria, Queensland); HT – ANIC. parvisetosus Ochi, Kon and Bai, 2010: 131; TT; China (Sichuan); HTm + PTs – CASB. pastillarius (Blanchard, 1846: 187); OO; Argentina, Bolivia, Chile; HT – MNHN? patagonicus (Blanchard, 1846: 186); Pn; Argentina, s. Brazil; HT – MNHN? patriciae Pittino, 1987: 386; Pn; Argentina, Uruguay; HTm + PTm – RPMI. pauliani (Haaf, 1954a: 385); OA; China (Henan, Hong Kong, Hubei, sw. China), India (Sikkim, West Bengal), Nepal, Sri Lanka, Cambodja, n. Vietnam, Annam, Indonesia (Java); HTm – NHMV, PTs – BMNH. —pectinatus (Pallas, 1781: 10), as Scarabaeus; type lost; = horridus (syn. by Harold 1872: 36). pedestris (Harold, 1872: 128); Pn; Argentina; HT – MNHN?; syn. denticulatus Blanchard (n. praeocc.). pellosomus Scholtz, 1986b: 27; OO; Australia (New South Wales, Victoria); HT – ANIC, PTs – BMNH. penicillatus Fåhraeus, 1857: 383; TP; RSA; LTm – NHRS (des. by Scholtz 1980a: 81). perhispidus (Blackburn, 1904: 296); OO; Australia (all states except Tasmania); HT – BMNH. perlatus (Goeze, 1777: 74), as Scarabaeus; TT; Europe, Algeria; syn. chevrolati, sabulosus Olivier, subterraneus. CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 15 perrieri Fairmaire, 1899: 519; TT; Madagascar; HTm – MNHN; not considered by Scholtz (1986a: 361), but compared by Haaf (1953: 342) to T. (T.) pusillus and therefore hereby placed in Trox s. str. rather than in subgenus Phoberus (see also Pittino 2010: 80). perrisii Fairmaire, 1868: 482; TT; Europe, Turkey, Algeria, Morocco; syn. haroldi, nidicola. persuberosus (Vaurie, 1962: 145); OO; Argentina, Bolivia, Brazil, Paraguay, Peru, Uruguay; HTm + PTs – SASP and other institutions. peruanus (Erichson, 1847: 111); Pn; Argentina, Bolivia, Chile, Peru; LTm – MNHB. —philbyi (Paulian, 1948: 143); HTm – MNHN; = niloticus desertorum (syn. by Haaf 1954a: 347). pilularius (Germar, 1824: 113); Pn; Argentina, Bolivia, Brazil, Chile, Paraguay, Uruguay; HT – MLUH? —pinguis Faldermann, 1835b: 369; = morticinii. placosalinus Ren, 2003: 112; TT; China (Xinjiang); HT – CASB? planicollis Haaf, 1953: 337; TP; Lesotho, Namibia, RSA (Cape), Zimbabwe; HTm – NHMB. —platycyphus (Loomis, 1922: 133); HT – USNM; = umbonatus (syn. by Robinson 1940: 150). plicatus Robinson, 1940: 153; TT; USA (Arizona, New Mexico, Texas), Mexico (most states); HT – ANSP. ponderosus (Péringuey, 1901: 467); OA; Ethiopia, Mozambique, e. RSA, Tanzania, Zimbabwe; type lost. —porcatus Say, 1825: 193; = unistriatus (syn. by Harold 1872: 174). poringensis Ochi, Kon and Kawahara, 2005: 65; TT; Borneo (Sabah), Java; HTf – UMBS; male described by Pittino and Kawai (2006: 7). principalis (Haaf, 1954a: 337), as ssp. of guttalis (elevated to species by Scholtz 1980a: 43); OA; n. Cameroon, Chad, e. Nigeria, Sudan; HTm – NHMB. procerus (Harold, 1872: 127); OA; Arabia, n. Burkina Faso, Chad, Egypt, Ethiopia, c. India, Iran, Mali, Niger, ne. Nigeria, Pakistan, Senegal Somalia, Sudan; LT – MNHN (des. by Scholtz 1980a: 35). punctatus (Germar, 1824: 113); OO; USA (w. states from Montana, Utah and California e. to the Mississippi River; also Kentucky, Ohio, Mississippi), Mexico (Aguascalientes, Baja, Chihuahua, Durango, Guanajuato, Nuevo León, Sonora); syn. integer, morsus. —punctatus LeConte, 1854: 215; HT – MCZH? [not found by Vaurie (1955: 61)]; = suberosus (syn. by Harold 1872: 120). puncticollis Haaf, 1953: 330; TP; Arab Emirates, Yemen; HTm – ZSBS, PTs – MNHN; Pittino 2006: 81 lists it in Trox s. str. pusillus Péringuey, 1908: 634; TT; Angola, Benin, Congo (DRC), Gabon, Ghana, Guinea, Ivory Coast, Liberia, Sierra Leone, Tanzania, Uganda, Zaire, Zambia, Zimbabwe; LT – SAMC (des. by Scholtz 1980a: 74); syn. opacus, pusio, urbani. —pusio Kolbe, 1914: 288; LT – MNHB; = pusillus (syn. and LT des. by Scholtz 1980a: 74). —pustulatus LeConte, 1854: 215; HT – MCZH; = monachus (syn. by Harold 1872: 116). quadricostatus Scholtz, 1980, see ssp. of natalensis. quadridens (Blackburn, 1892: 38); OO; Australia (all states except Tasmania); LT – BMNH (des. by Scholtz 1986b: 41). quadrimaculatus Ballion, 1871: 338; TT; Israel, Iran, Kazakhstan, Kyrgyzstan, Tajikistan, Uzbekistan; type – ZMAS?, ZMUM? quadrinodosus (Haaf, 1954b: 723); OO; Australia (Capital Ter., New South Wales, Queensland, Victoria); HT – MNHB, PTs – BMNH. radula (Erichson, 1843: 235); OA; Angola, Botswana, Kenya, Mozambique, Namibia, e. RSA, Tanzania, Zimbabwe; LTm – MNHB (des. by Scholtz 1979a: 190); syn. variolosus. —raduloides (Kolbe, 1904: 292); LT – MNHB; = tuberosus (syn. and LT des. Scholtz 1980a: 44). —ranavalo (Paulian, 1937: 134); HT – BMNH; = sallei (syn. by Vaurie 1962: 130). ranotsaraensis Pittino, 2010: 77; MT; sw. Madagascar; HTf – MNHN; no other material. regalis (Haaf, 1954b: 704); OO; “Australia”; HTm – MNHN; known only from HT. —regularis (Harold, 1868: 85); = costatus (syn. by Harold 1872: 115). reiterorum Král and Kubán, 2012: 148; OA; Socotra I.; HT + AT + PTs – NMPC. rhyparoides Harold, 1872: 32; TT; Lesotho, RSA, St. Helena I. (intro.); LT – MNHN (des. by Scholtz 1980a: 73); syn. whiteheadi. 16 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK rimulosus (Haaf, 1957a: 693); OA; India (“Vorder-Indien: Coromandel”); HTm – NHMB. robinsoni Vaurie, 1955: 47; TT; Canada (Alberta, Manitoba, Saskatchewan), USA (c. states s. to Texas; also Colorado, Illinois, Michigan, Minnesota, Wisconsin); HT + AT + PTs – AMNH, PTs also at other institutions. rotundulus (Haaf, 1957b: 117); OO; Australia (Northern Ter., Western Au.); HT – MVMA rowei Scholtz 1980, see ssp. of fascicularis. rubricans (Robinson, 1946: 57); OO; USA (se. Texas), Mexico (widespread), Guatemala (w. coast), Nicaragua (w. coast); HTm – CASC. rudebecki Haaf, 1958a: 474; TT; Lesotho, RSA (sw. Cape, w. Natal); HTm – MZLU. rusticus (Fåhraeus, 1857: 379); OA; Namibia, e. RSA, Zimbabwe; LT – NHRS (des. Scholtz 1979a: 191); syn. damarinus. sabulosus sabulosus (Linné, 1758: 350), as Scarabaeus; TT; Europe, Siberia, Kazakhstan, Mongolia, Korean peninsula, Turkey, Syria; type – ZMUU?; syn. femoratus. sabulosus fujiokai Ochi, 2000: 43; Japan (Honshu). —sabulosus Olivier, 1789: no. 4: 8; = perlatus. —sabulosus Preyssler, 1790: 48; = hispidus. salebrosus (MacLeay, 1871: 186); OO; Australia (Northern Ter., Queensland, Western AU.); LT – AMSA (des. by Scholtz 1986b: 32); syn. vitreomaculatus. sallei (Harold, 1872: 138); Pn; Bolivia, Ecuador, Peru; HT – MNHN; syn. ranavalo. satorui Kawai, 2006: 1; OA; RSA (Cape), Ivory Coast; HTm – NSMT. scaber (Linné, 1767: 573); TT; Europe, Siberia, n. Africa, Canary Is., Canada, USA, Argentina, Chile (recorded but not established in Chile, see Diéguez 2008), Australia, Borneo (Sabah); type – ZMUU?; syn. arenarius, arenosus, barbosus, hispidus Paykull, niponensis, trisulcatus. scabrosus (Beauvois, 1805: 175); OO; Canada (Ontario), USA (from New York to Florida and w. to Nebraska, Colorado, Kansas, Oklahoma and Texas). schaborti Scholtz, 1980, see ssp. of natalensis. scutellaris (Say, 1823: 238); OO; USA (Arizona, Colorado, Kansas, New Mexico, ne. Utah, Texas), Mexico (Chihuahua, Coahuila, D.F., Durango, Morelos, Puebla, Sonora); LT? – MCZH. semicostatus (MacLeay, 1871: 186); OO; Australia (Capital Ter., New South Wales, Queensland, Western AU.); LT – ANIC (des. by Scholtz 1986b: 31); syn. asperrimus. —semmelinki (Lansberge, 1887: 200); = squamosus (syn. by Scholtz 1986b: 53). senegalensis (Scholtz, 1983: 205); OA; Senegal; HTm – TMSA, PTf – UPSA (types only). —serrulatus Beauvois, 1805: 176; = tuberculatus DeGeer (syn. by Harold 1872: 155). setifer Waterhouse, 1875: 98; TT; Japan, South Korea; LT + PLT – BMNH; syn. aino. setosipennis (Blackburn, 1904: 292); OO; Australia (New South Wales, Northern Ter., Queensland, South Au.); LT – BMNH (des. by Scholtz 1986b: 23). —setulosus Kolbe, 1891: 23; HTm – MNHB; = squamiger (syn. by Haaf 1953: 326). seymourensis (Mutchler, 1925: 229); Pn; s. + c. Galapagos; HT – AMNH. sibericus Nikolaev, 2007: 104f, 215, 220; Trox inc. sed.; fossil (Early Cretaceous), Siberia. —silphoides Thunberg, 1787: 38; type – ZMUU?; = horridus (syn. by Harold 1872: 36). simi Robinson, 1940; ssp. of spinulosus. sonorae LeConte, 1854: 211; TT; Canada (Alberta, Northwest Ter., Saskatchewan), USA (e. to South Dakota, Nebraska, Kansas and Texas; also California, Louisiana, Missouri), Mexico (Chihuahua, Durango, Nuevo León, Sonora, Zacatecas); HT – MCZH; syn. alternans LeConte, lecontei. sordidatus Balthasar, 1936: 452, as ssp. of granulipennis; TT; Bulgaria, Croatia, Greece, Syria, Turkey; HTm[ST?] – NMPC (elevated to species by Pittino 1983: 277). sordidus LeConte, 1854: 211; TT; Canada (Ontario, Quebec), USA (most e. states and w. to Kansas and Texas); HT [=LT] – MCZH. spatulatus (Vaurie, 1962: 151); OO; Argentina; HTm – NHMB, PTs – CMNO. —speculifer (Heller, 1914: 317); = squamosus (syn. by Scholtz 1986b: 53). spinulosus spinulosus Robinson, 1940: 156; TT; Canada (Ontario), USA (from Michigan to Tennessee and w. to Nebraska; also Arkansas, Louisiana); HT – ANSP. CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 17 spinulosus dentibius Robinson, 1940: 156, as sp. (reduced to ssp. by Vaurie 1955; 36); USA (Arizona, New Mexico, w. Texas), Mexico (Baja, Chihuahua, Coahuila, Durango, Michoacán, Morelos, Nuevo León); HT – ANSP; intergrades with spinulosus s. str. in sc. Texas. spinulosus simi Robinson, 1940: 157, as species (reduced to ssp. by Vaurie 1955: 36); USA (e. states from Pennsylvania and New Jersey s. to Virginia); HT – ANSP. squalidus (Olivier, 1789: 12); OA; Afrotropical: continent-wide and Madagascar; LTm – TMSA (des. Scholtz 1979a: 187); syn. gonoderus, incultus. squamiger Roth, 1851: 133; TP; Burundi, Cameroon, Ethiopia, Kenya, RSA, Rwanda, Somalia, Tanzania, Uganda, Zaire, Zimbabwe [listed also from the Arab Emirates and Yemen, see Pittino 2006: 81]; LTf – ZSMC (des. by Scholtz 1980a: 90); syn. angulatus, setulosus. squamosus (MacLeay, 1871: 186); OO; e. Australia, e. Papua New Guinea; LT – ANIC (des. by Scholtz 1986b: 53); syn. semmelinki, speculifer. stellatus (Harold, 1872: 92); OO; Australia (Western Au.); HT – NHMV. strandi Balthasar, 1936: 452; TT; Algeria, Libya, Morocco, Tunisia; HT[ST?] – NMPC. striatus Melsheimer, 1846: 137; TT; Canada (Ontario), USA (Arkansas, Illinois, New Jersey, Pennsylvania); HT – MCZH? (not found by Vaurie 1955: 32). strigosus Haaf, 1953: 319; TT; RSA (e. Cape, Free State, se. Transvaal); HTm – NHMB, PTs – BMNH. strzeleckensis (Blackburn, 1895: 33); OO; Australia (all states except Tasmania); LT – BMNH (des. by Scholtz 1986b: 24). subcarinatus (MacLeay, 1864: 128); OO; Australia (all states except Tasmania), Papua New Guinea; LT – ANIC (des. by Scholtz 1986b: 51); syn. fenestratus, nodicollis? suberosus (Fabricius, 1775: 31); OO; New World, Australia, Oceania; HTm – BMNH; syn. alternatus, crenatus, denticulatus, gibbus, manilensis, nobilis, novaecaledoniae, ovatus, punctatus LeConte, torresalei, tricolor, triestinae, tuberosus Castelnau. —subterraneus (Geoffroy, 1785: 8), as Scarabaeus; = perlatus. sugayai Masumoto and Kiushi, 1995: 206; TT; s. Japan. sulcatus Thunberg, 1787: 38; TP; Namibia, RSA, Zimbabwe; type(s) – ZMUU; syn. instabilis. —suturalis LeConte 1854: 214; LTm – MCZH (des. by Robinson 1940: 150); = texanus (syn. by Vaurie 1955: 75). taiwanus Masumoto, Ochi and Li, 2005: 233; TT; c. Taiwan. talpa Fåhraeus, 1857: 380; TP; RSA (e. Cape, Natal); HT – NHRS. tasmanicus (Blackburn, 1904: 292); OO; Australia (Tasmania + Flinders I.); HT – BMNH. tatei (Blackburn 1892: 37); OO; Australia (New South Wales, South Au., Western Au.); LT – BMNH (des. by Scholtz 1986b: 11). tenebrosus (Harold, 1872: 130); Pn; Ecuador; HTm – MNHN. terrestris Say, 1825: 192; TT; USA (Pennsylvania and Atlantic coast states from Massachusetts s. to Mississippi and Florida); types destroyed. tesselatus LeConte, 1854: 216; OO; USA (Arizona, Kansas, New Mexico, Texas), Mexico (Baja, Guerrero, Morelos, Sinaloa, Sonora); LTm – MCZH (des. by Robinson 1940: 151); syn. nigrescens. testudo (Arrow, 1927: 466); OA; India, Pakistan; HTm – BMNH. texanus LeConte, 1854: 214; OO; USA (New Mexico, s. Texas), Mexico (Nuevo León, Tamaulipas); LTf – MCZH (des. by Robinson 1940: 150). tibialis Masumoto, Ochi and Li, 2005: 235; TT; Taiwan. tomentosus (Robinson, 1941a: 135); OO; Mexico (Nayarit, Oaxaca, Sinaloa, Sonora); HTm – ANSP. —torresalei (Báguena Corella, 1959: 212); = suberosus (syn. by Scholtz 1982: 14). transversus Reiche, 1856: 405; TT; Greece; LT – HNHM (des. by Pittino 2006: 28); syn. demaisoni, graecus. —tricolor (Blackburn, 1904: 296); type – BMNH; = suberosus (syn. by Arrow 1927: 466). —triestinae Pittino, 1987: 378; HTm – MNHN; = suberosus (syn. by Scholtz 1990b: 1407). trilobus (Haaf, 1954b: 727), as ssp. of candidus; OO; Australia (all states except Western Au. and Tasmania), Papua New Guinea; HT + PTs – BMNH. —trisulcatus Curtis, 1845: 446; HT – BMNH; = scaber (syn. by Harold 1872: 183). 18 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK tuberculatus (DeGeer, 1774: 318), as Scarabaeus; TT; USA (from Massachusetts s. to extreme nw. Florida and w. to e. Texas, e. Kansas and Nebraska; also Arizona, New Mexico, Wyoming); syn. canaliculatus, serrulatus. —tuberculatus (Beauvois, 1805: 175); = monachus (syn. by Harold 1872: 116). —tuberosus (Castelnau, 1840: 107); = suberosus (misspelling, see Vaurie 1955: 60). tuberosus (Klug, 1855: 657); OA; Botswana, Ethiopia, Kenya, Mozambique, e. RSA, Tanzania, Uganda, Zaire; LTm – MNHB (des. by Scholtz 1980a: 44); syn. raduloides. tytus (Robinson, 1941b: 228); OO; USA (Arizona, Georgia, Oklahoma, Pennsylvania), Cuba; HTm – ANSP, PT – BMNH; known only from nests of barn owl (Tyto alba pratincola). uenoi uenoi Nomura, 1961: 70; TT; Japan (Tokuno-shima Is.); HTm – NSMT. uenoi matsumurai Miyake and Yamaya, 1995: 31; Japan (Okinawa-shoto). umbonatus LeConte, 1854: 214; OO; USA (Arizona, New Mexico, Texas, Utah), Mexico (Chihuahua, Coahuila, Nuevo León, Tamaulipas); LTm – MCZH (des. by Robinson 1940: 150); syn. confluentus, platycyphus. —undulatus Zoubkoff, 1829: 158; = cadaverinus cadaverinus. unguicularis (Haaf, 1954a: 369); OA; Mozambique, ne. RSA, e. Zimbabwe; HTf – NHMB, PTs – BMNH, FMNH and other institutions. unistriatus Beauvois, 1805: 175; TT; Canada (all provinces except British Columbia),USA (s. to Colorado in the west and to North Carolina in the east; also sw. Arkansas, Florida, Louisiana, Texas); syn. porcatus. —urbani Miksic, 1958: 15; type lost; = pusillus (syn. by Scholtz 1980a: 74). ussuriensis Balthasar, 1931: 129; TT; Siberia, Mongolia, China (Heilongjiang), Korean peninsula; STs(2) – NMPC. varicosus (Erichson, 1843: 235); OA; w. Angola; LTm – ISNB (des. by Scholtz 1980a: 63). variolatus Melsheimer, 1846: 138; TT; Canada (New Brunswick, Nova Scotia, Ontario, Quebec), USA (e. states w. to Kansas, Oklahoma and Texas; also Iowa); HT – MCZH; syn. erinaceus, variolatus monticola. —variolosus (Fåhraeus, 1857: 378); LT – NHRS (des. Scholtz 1980a: 47), PLT(1) – BMNH; = radula (syn. by Harold 1872: 68). —velutinus (Blackburn, 1892: 40); HT – BMNH; = costatus (syn. by Haaf 1954b: 734). verrucosus (Reiche, 1856: 404); OA; e. Africa incl. Egypt, Arabian peninsula, Iran Iraq; LTm – MNHN (des. by Pittino 2006: 28), “one type” – BMNH; syn. nomadicus; considered a synonym of denticulatus (Olivier) by Harold (1872), Arrow (1927), Haaf (1954a) and Scholtz (1980a, 1982), revalidated by Pittino (2006: 28). villosus (Haaf, 1954b: 728); OO; Australia (Northern Ter., Queensland, Western Au.); HT – MNHB, PTs – BMNH. vimmeri Balthasar, 1931: 130; TT; e. Siberia, Mongolia, China (Beijing), South Korea; HT[ST?] – NMPC; revived from synonymy of T. ineptus by Pittino (2006: 27). —vitreomaculatus (MacLeay, 1888: 909); = salebrosus (syn. by Blackburn 1904: 284). vladislavi Kawai, 2009: 73; OO; Australia (n. Queensland); HT – QMBA, AT + PTs – VMPC? —whiteheadi Wollaston, 1877: 61; HT – BMNH; = rhyparoides (syn. by Haaf 1953: 340). wittei (Haaf, 1955a: 47), as ssp. of guttalis (elevated to species by Scholtz 1980a: 43); OA; Angola, Namibia, Tanzania, s. Zaire, Zambia, Zimbabwe; HTm – ISNB. yamayai Nakane, 1983: 139; TT; Japan (Ryukyus: Iriomote I.); HTm – TNCJ. yangi Masumoto, Ochi and Li, 2005: 238; TT; n. Taiwan. youngai Struempher and Scholtz, 2011: 337; TT; RSA (possibly extinct). zoufali Balthasar, 1931: 131; TT; e. Siberia, China (Beijing, Hubei), North Korea, Oriental; HT[ST?] – NMPC. zumpti (Haaf, 1957a: 692); OA; Botswana, Kenya, Namibia, e. RSA, Zimbabwe; HTm + PTs – NHMB. CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 19 Trox (Trox) acanthinus Harold, Nt aequalis Say, Na-Nt L affinis Robinson, Na alpigenus Zhang, Pa aproximans Escalera, Pa atrox LeConte, Na-Nt L boucomonti Paulian, Pa-Or brahminus Pittino, Pa-Or cadaverinus Illiger, Pa cambeforti Pittino, Pa cambodjanus Pittino, Or capensis Scholtz, Af L capillaris Say, Na clathratus Reiche, Pa confluens Wollaston, Pa contractus Robinson, Na cotodognanensis Compte, Pa cribrum Gené, Pa L dhaulagiri Paulus, Pa eversmanni Krynicki, Pa eximius Faldermann, Pa fabricii Reiche, Pa fascicularis Wiedemann, Af L fascifer LeConte, Na L floridanus Howden and Vaurie, Na formosanus Nomura, Pa foveicollis Harold, Na L frontera Vaurie, Na L gansuensis Ren, Pa gemmulatus Horn, Na-Nt granulipennis Fairmaire, Pa-Af hamatus Robinson, Na hispanicus Harold, Pa hispidus (Pontoppidan), Pa L horiguchii Ochi and Kawahara, Pa inadai Ochi et al., Pa ineptus Balthasar, Pa insularis Chevrolat, Nt iranicus Petrovitz, Pa jeanae Scholtz et al., Or klapperichi Pittino, Pa koreanus Kim, Pa kyotensis Ochi and Kawahara, Pa lama Pittino, Pa laticollis LeConte, Na leonardii Pittino, Pa litoralis Pittino, Pa Lists of valid species by genera and subgenera Subspecies, nomina dubia and fossil species indeterminate to subgenus omitted. Limits of biogeographic regions after Löbl and Smetana (2006: 12, map 1). Af–Afrotropical, Au–Australian, Md–Madagascan, Na–Nearctic, Nt–Neotropical, Or–Oriental, Pa–Palearctic. L–larva known. mandli Balthasar, Pa martini Reitter, Pa matsudai Ochi and Hori, Pa mitis Balthasar, Pa mixtus Harold, Pa morticinii (Pallas), Pa mutsuensis Nomura, Pa nanniscus Péringuey, Af nasutus Harold, Af natalensis Haaf, Af necopinus Scholtz, Af niger Rossi, Pa niponensis Lewis, Pa-Or nodulosus Harold, Pa nohirai Nakane, Pa opacotuberculatus Motschulsky, Pa parvisetosus Ochi et al., Pa perlatus (Goeze), Pa L perrieri Fairmaire, Md perrisii Fairmaire, Pa placosalinus Ren, Pa plicatus Robinson, Na-Nt L puncticollis Haaf, Af pusillus Péringuey, Af quadrimaculatus Ballion, Pa rhyparoides Harold, Af L robinsoni Vaurie, Na L rudebecki Haaf, Af sabulosus (Linné), Pa L scaber (Linné), subcosmopolitan L setifer Waterhouse, Pa sonorae LeConte, Na-Nt L sordidatus Balthasar, Pa sordidus LeConte, Na L spinulosus Robinson, Na-Nt L squamiger Roth, Pa strandi Balthasar, Pa striatus Melsheimer, Na strigosus Haaf, Af sugayai Masumoto and Kiushi, Pa terrestris Say, Na transversus Reiche, Pa tuberculatus (DeGeer), Na L uenoi Nomura, Pa unistriatus Beauvois, Na L ussuriensis Balthasar, Pa variolatus Melsheimer, Na-Nt L vimmeri Balthasar, Pa 20 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK Trox (Trox), continued yamayai Nakane, Pa youngai Struempher and Scholtz, Af zoufali Balthasar, Pa-Or Trox (Madagatrox) [monobasic] ranotsaraensis (Pittino), Md Trox (Phoberus) [all Af] aculeatus Harold arcuatus Haaf braacki Scholtz brincki Haaf caffer Harold consimilis Haaf L cyrtus Haaf gunki Scholtz horridus Fabricius levis Haaf luridus Fabricius miliarius Gmelin montanus Kolbe nama Kolbe nigrociliatus Kolbe penicillatus Fåhraeus planicollis Haaf squamiger Roth L sulcatus Thunberg talpa Fåhraeus Omorgus (Haroldomorgus) [monobasic] badeni (Harold), Nt Omorgus (Omorgus) alatus (MacLeay), Au alius Scholtz, Au alternans (MacLeay), Au amictus (Haaf), Au aphanocephalus Scholtz, Au asper LeConte, Na-Nt L augustae (Blackburn), Au australasiae (Erichson), Au batesi (Harold), Nt birmanicus (Arrow), Or borrei (Harold), Nt brucki (Harold), Au candezei (Harold), Nt candidus (Harold), Au capillaceus Scholtz, Nt carinatus (Loomie), Na-Nt L carinicollis Scholtz, Au ciliatus (Blanchard), Nt costatus (Wiedemann), Au-Or L crotchi (Harold), Au curvipes (Harold), Au demarzi (Haaf), Au dilaticollis (MacLeay), Au dohrni (Harold), Au elderi (Blackburn), Au elongatus (Haaf), Au euclensis (Blackburn), Au eyrensis (Blackburn), Au fuliginosus (Robinson), Na-Nt L gigas (Harold), Au granuliceps (Haaf), Au howdenorum Scholtz, Au howelli (Howden and Vaurie), Na-Nt indigenus Scholtz, Nt inflatus (Loomie), Na-Nt L insignicollis (Blackburn), Au loxus (Vaurie), Nt lucidus Pittino, Nt mariae Scholtz, Au mariettae Scholtz, Au marshalli (Haaf), Au mentitor (Blackburn), Au mictlensis Deloya, Nt mollis (Arrow), Or monachus (Herbst), Na-Nt L monteithi Scholtz, Au nanningensis Pittino, Pa nigroscobinus Scholtz, Au nocheles Scholtz, Nt nodicollis (MacLeay), Au nodosus (Robinson), Na-Nt ovalis (Haaf), Au parvicollis Scholtz, Au pastillarius (Blanchard), Nt pellosomus Scholtz, Au perhispidus (Blackburn), Au persuberosus (Vaurie), Nt L punctatus (Germar), Na-Nt L quadridens (Blackburn), Au quadrinodosus (Haaf), Au regalis (Haaf), Au rotundulus (Haaf), Au rubricans (Robinson), Na-Nt L salebrosus (MacLeay), Au scabrosus (Beauvois), Na-Nt scutellaris (Say), Na-Nt L semicostatus (MacLeay), Au setosipennis (Blackburn), Au spatulatus (Vaurie), Nt squamosus (MacLeay), Au stellatus (Harold), Au strzeleckensis (Blackburn), Au subcarinatus (MacLeay), Au suberosus (Fabricius), Na-Nt-Au L CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 21 Omorgus (Omorgus), continued tasmanicus (Blackburn), Au tatei (Blackburn), Au tesselatus LeConte, Na-Nt texanus LeConte, Na-Nt L tomentosus (Robinson), Nt trilobus (Haaf), Au tytus (Robinson), Na-Nt umbonatus LeConte, Na-Nt villosus (Haaf), Au vladislavi Kawai, Au Omorgus (Afromorgus) acinus (Scholtz), Af amitinus (Kolbe), Af asperulatus (Harold), Af L baccatus (Gerstaecker), Af borgognoi (Marchand), Af chinensis (Boheman), Pa-Or consanguineus (Péringuey), Af denticulatus (Olivier), Af discedens (Haaf), Af elevatus (Harold), Af endroedyi (Scholtz), Af expansus (Arrow), Af foveolatus (Boheman), Af frater Pittino, Pa-Or freyi (Haaf), Af L funestus (Lansberge), Af gemmatus (Olivier), Pa-Af genieri Scholtz, Af granulatus (Herbst), Pa-Or guttalis (Haaf), Pa-Af haagi (Harold), Pa-Or inclusus (Walker), Pa-Or indicus (Harold), Or inermis Pittino, Or insignis (Haaf), Af italicus (Reiche), Pa-Or lindemannae (Petrovitz), Af lobicollis (Arrow), Or lugubris (Haaf), Af maindroni Pittino, Pa melancholicus (Fåhraeus), Pa-Af-Md L mutabilis (Haaf), Af niloticus (Harold), Pa-Af obesus (Scholtz), Af omacanthus (Harold), Pa-Or pauliani (Haaf), Pa-Or ponderosus (Péringuey), Af principalis (Haaf), Af procerus (Harold), Af-Pa-Or L radula (Erichson), Af L reiterorum Král and Kubán, Pa-Af (Socotra) rimulosus (Haaf), Or rusticus (Fåhraeus), Af satorui Kawai, Af senegalensis (Scholtz), Af squalidus (Olivier), Af-Md L testudo (Arrow), Pa-Or tuberosus (Klug), Af unguicularis (Haaf), Af varicosus (Erichson), Af verrucosus (Reiche and Saulcy), Pa-Af wittei (Haaf), Af zumpti (Haaf), Af Polynoncus [all Nt] aeger (Guérin) aricensis (Gutiérrez) L bifurcatus (Vaurie) brasiliensis (Vaurie) brevicollis (Eschscholtz) bullatus (Curtis) burmeisteri Pittino chilensis (Harold) diffluens (Vaurie) ecuadorensis (Vaurie) erugatus Scholtz galapagoensis (Van Dyke) gemmifer (Blanchard) gemmingeri (Harold) gibberosus Scholtz gordoni (Steiner) guttifer (Harold) haafi (Vaurie) hemisphaericus (Burmeister) juglans (Ratcliffe) longitarsis (Harold) mirabilis Pittino neuquen (Vaurie) pampeanus (Burmeister) parafurcatus Pittino patagonicus (Blanchard) patriciae Pittino pedestris (Harold) peruanus (Erichson) pilularius (Germar) sallei (Harold) seymourensis (Mutchler) L tenebrosus (Harold) 22 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK Acknowledgments I am grateful to Philip D. Perkins (MCZH) for commenting on the putative types of Say, Bushra M. Hussaini (AMNH) for information on the type repository of Trox antiquus Wickham, and Maxwell V. L. Barclay (BMNH) for pagination of MacLeay 1826. W. David Edmonds (Marfa, Texas) and Ales Bezdek (Czech Academy of Sciences, Ceske Budejovice) kindly helped with literature and their critical comments much improved the manuscript. Literature cited Ádám, L. 1994. A new species of the genus Trox. Folia Entomologica Hungarica 55: 37-39. Arnett, R. H. Jr., G. A. Samuelson, and G. M. Nishida. 1993. The insect and spider collections of the world. Second edition. Sandhill Crane Press; Gainesville, FL. 310 p. Arrow, G. J. 1900. On a collection of insects and arachnids made in 1897 by Mr. C. K. A. Peel in Somaliland. Proceedings of the Zoological Society of London 1900: 4-63. Arrow, G. J. 1903. On the laparostict Coleoptera of Granada and St. Vincent (W. Indies). Transactions of the Entomological Society of London 1903: 509-520. Arrow, G. J. 1909. On the characters and relationships of the less known groups of lamellicorn Coleoptera. Transactions of the Entomological Society of London 1909: 479-507. Arrow, G. J. 1912. Trogidae. Coleopterorum Catalogus 19(43): 52-66. Arrow, G. J. 1927. A note on the coleopterous genus Trox, with description of a few new Asiatic species. Annals and Magazine of Natural History (9) 19: 465-468. Báguena, L. 1959. Cuarro novedades y un comentario sobre coleópteros de España. Eos 35: 209-214. Baker, C. W. 1968. Larval taxonomy of the Troginae in North America with notes on biologies and life histories (Coleoptera: Scarabaeidae). Bulletin of the U. S. National Museum 279: 1-79. Ballion, E. 1871. Eine Centurie neuer Käfer aus der Fauna des russischen Reiches. Bulletin de la Société Impériale des Naturalistes de Moscou 43(2): 320-353. Balthasar, V. 1931. Sechs neue Trox-Arten aus dem Ussurigebiet und Transbaikalien. Entomologische Blätter 27: 128-134. Balthasar, V. 1933. Zwei neue Formen von Scarabaeiden. Entomologische Nachrichtenblatt 7: 81-84. Balthasar, V. 1936. Monographie der Subfam. Troginae der palaearktischen Region. In Festschrift für Prof. Dr. Embrik Strand, Vol. l: 407-459. Riga. Balthasar, V. 1939. Neue Arten der coprophagen Scarabaeiden aus dem Museo Zoologico della Università di Firenze. Redia 25: 1-36. Balthasar, V. 1966. Neue Gattungen und Arten der Scarabaeoidea der australischen und neotropischen Regionen. Entomologische Blätter 62: 177-185. Beauvois, A. M. F. J. Palisot de 1805. Insectes recueillis en Afrique et en Amérique, dans les Royaumes d’Oware et de Benin, à Saint-Dominique et dans les États-Unis, pendant les années 1786-1797. Levraut; Paris. 276 p., 90 pls. [1805-1821, issued in parts]. Berg, F. G. C. 1884. Coléoptères de Buenos Aires. Bulletin des Séances de la Société Entomologique de France (6) 4: 48-111. Blackburn, T. 1892. Further notes on Australian Coleoptera, with descriptions of new genera and species, 11. Transactions of the Royal Society of South Australia 15: 20-73. Blackburn, T. 1895. Further notes on Australian Coleoptera, with descriptions of new genera and species, 17. Transactions of the Royal Society of South Australia 19: 27-60. Blackburn, T. 1896. Coleoptera (exclusive of Carabidae). Report of the Horn Expedition to central Australia, pt. 2, Zoology. Transactions of the Royal Society of South Australia 20: 254-308. Blackburn, T. 1904. Further notes on Australian Coleoptera, with descriptions of new genera and species, 34. Transactions of the Royal Society of South Australia 28: 281-297. Blanchard, E. 1846. Insectes de l’Amérique recueillis par Alcide d’Orbigny. In d’Orbigny, A. Voyage dans l’Amérique méridionale, vol. 6, pt. 2. Bertrand, Paris. 222 p. Boheman, C. H. 1858. Coleoptera. Species novas descripsit. In: Kongliga Svenska Fregatten Eugenies resa omkring Jorden ... Zoologi, I, Insecta. Norstedt; Stockholm. 112 p. CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 23 Boheman, C. H. 1860. Coleoptera samlade J. A. Wahlberg i Syd-Vestra Africa. Öfversigt af Kongliga Svenska Vetenskaps-Akademiens Förhandlingar 17: 107-120. Bonnaire, A. 1881. Coléoptères de Fontainbleau et de l’Île de Ré. Bulletin de la Société Entomologique de France 6: 62-68. Browne, D. J., and C. H. Scholtz. 1995. Phylogeny of the families of Scarabaeoidea (Coleoptera) based on characters of the hindwing articulation, hindwing base and wing venation. Systematic Entomology 20: 145-173. Browne, D. J., and C. H. Scholtz. 1996. Morphology of the hind wing articulation and wing base of the Scarabaeoidea (Coleoptera) with some phylogenetic implications. Bonner Zoologische Monographien 40: 1-200. Browne, D. J., and C. H. Scholtz. 1998. Evolution of the scarab hindwing articulation and wing base: a contribution toward the phylogeny of the Scarabaeidae (Scarabaeoidea: Coleoptera). Systematic Entomology 23: 307-326. Browne, D. J., and C. H. Scholtz. 1999. A phylogeny of the families of Scarabaeoidea (Coleoptera). Systematic Entomology 24: 51-84. Browne, D. J., C. H. Scholtz, and J. Kukalová-Peck. 1993. Phylogenetic significance of wing characters in the Trogidae (Coleoptera: Scarabaeoidea). African Entomology 1(2): 195-206. Burmeister, H. C. C. 1876. Die argentinischen Arten der Gattung Trox F. Stettiner Entomologische Zeitung 37: 241-268. Castelnau, F. L. 1840. Histoire naturelle des insectes coléoptères. Tome deuxième. Histoire naturelle des animaux articulés, annelides, crustacés, arachnides, myriapodes et insectes. P. Duménil; Paris. 564 p. Chevrolat, L. A. A. 1864. Coléoptères de l’Île de Cuba, 5. Annales de la Société Entomologique de France 4(4): 405-418. Chevrolat, L. A. A. 1881. [Trox insularis Chevrolat = T. foveicollis Harold]. Annales de la Société Entomologique de France 6(1): 145. Curtis, J. 1845. Descriptions ... of insects collected by Captain P. P. King ... in the survey of the Straits of Magellan. Transactions of the Linnean Society of London 19: 441-475 [+ pl. 41]. DeGeer, C. 1774. Mémoires pour servir à l’histoire des Insectes. Tome quatrième. L’imprimerie Pierre Hesselberg; Stockholm. 456 p. Deloya, C. 1995. A new species of Omorgus (Omorgus) Erichson from Mexico. The Coleopterists Bulletin 49(2): 153-156. Deloya, C. 2003. Familia Trogidae. p. 125-133. In: M. Á. Morón. (ed.). Atlas de los escarabajos de México. Coleoptera: Lamellicornia, Vol. II. Familias Scarabaeidae, Trogidae, Passalidae y Lucanidae. Argania editio; Barcelona. 227 p. Diéguez, V. M. 2008. Present knowledge of the Trogidae of Chile (Coleoptera: Scarabaeoidea). Revista Chilena de Entomología 34: 11-28. Erichson, W. F. 1842. Beitrag zur Fauna von Vandimensland mit besonderer Rücksicht auf die geographische Verbreitung der Insecten. Archiv für Naturgeschichte 8: 85-287, pls. 4-5. Erichson, W. F. 1843. Beitrag zur Insecten-Fauna von Angola im besondere Beziehung zur geographischen Verbreitung der Insecten in Afrika. Archiv für Naturgeschichte 9: 199- 267. Erichson, W. F. 1847. Conspectus coleopterorum quae in Republica Peruana observata sunt. Archiv für Naturgeschichte 13(1): 67-185. Erichson, W. F. 1848 (1845-1848). Scaphidiidae – Scarabaeidae. In: Naturgeschichte der Insecten Deutschlands. Erste Abtheilung. Coleoptera. Dritter Band. Nicolaische Buchhandlung; Berlin. 968 p. Escalera, M. M. 1914. Los Coleópteros de Marruecas. Trabajos des Museo de Ciencias Naturales, Madrid, Serie Zoologica 11: 1-553. Eschscholtz, J. F. G. 1822. Descriptions d’Insectes nouveaux. Entomographien. Erste Lieferung. G. Reimer; Berlin. 128 p. Fabricius, J. C. 1775. Systema Entomologiae sistens Insectorum Classes, Ordines, Genera, Species adiectis Synonymis, Locis, Descriptionibus, Observationibus. Officina Libraria Kortii; Flensburgi et Lipsiae. 832 p. 24 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK Fabricius, J. C. 1781. Species insectorum exhibens eorum differentias specificas, synonyma auctorum, loca natalia, metamorphosis adiectis observationibus. Tomus I. Impensis C. E. Bohnii; Hamburgi et Kilonii. 494 p. Fabricius, J. C. 1787. Mantissa insectorum sistens eorum species nuper detectus adiectis characteribus genericis, differentiis specificis, emendationibus, descriptionibus. Tom. I. C. G. Proft; Hafniae. 348 p. Fabricius, J. C. 1792. Entomologia systematica emendata et aucta. Secundum classes, ordines, genera , species adiectis, synonymis, locis, observationibus, descriptionibus. Tom. I. C. G. Proft; Hafniae. 330 p. Fabricius, J. C. 1801. Systema Eleutheratorum secundum ordines, gnenera, species: adiectis synonymis, locis, observationibus, descriptionibus. Tom. I, II. Bibliopoli Academici Novi; Kiliae. I: 506 p., II: 687 p. Fåhraeus, O. J. 1857. Scarabaeidae et Lucanidae. In: C. H. Boheman, Insecta Caffrariae annis 1838-1845 a J. A. Wahlberg collecta amici auxilio suffultus descripsit. Pars II. Coleoptera. Officina Norstedtiana. 395 p. Fairmaire, L. 1852. Quelques Coléoptères nouveaux du midi de l’Europe et du nord de l’Afrique. Annales de la Société Entomologique de France (2) 10: 69-93. Fairmaire, L. 1868. Essai sur les Coléoptères de Barbarie. Sixième partie. Annales de la Société Entomologique de France (4) 8: 471-502. Fairmaire, L. 1871. Notes sur les Coléoptères recueillis par Charles Coquérel à Madagascar et sur les côtes d’Afrique (3). Annales de la Société Entomologique de France (5) 1: 29-78. Fairmaire, L. 1883. Notes sur quelques Coléoptères de Magellan et de Santa Cruz. Annales de la Société Entomologique de France (6) 3: 483-506. Fairmaire, L. 1893. Coléoptères des Îles Comores. Annales de la Société Entomologique de Belgique 37: 521-555. Fairmaire, L. 1899. Matériaux pour la faune coléoptèrologique de la région Malgache. Annales de la Société Entomologique de Belgique 43: 511-558. Fairmaire, L. 1901. Matériaux pour la faune coléoptèrologique de la région Malgache (11e-note). Revue d’Entomologie 20: 101-248. Faldermann, F. 1835a. Additamenta entomologica ad faunam rossicam in itineribus Jussu Imperatoris Augustissimi annis 1827-1831 a Cl. Ménétriés et Szovitz susceptis collecta, in lucem edita. Nouveaux Mémoires de la Société Impériale des Naturalistes de Moscou (2) 4: 1-310. Faldermann, F. 1835b. Coleopterorum ab illustrissimo Bungio in China boreali, Mongolia, et Montibus Altaicis collectorum, nec non ab ill. Turczaninoffio et Stchukino e provincia Irkutsk missorum illustrationes. Mémoires de l’Académie Impériale des Sciences de St. Pétersbourg (6) 3(1): 337-464. Felsche, C. 1901. Synonymische Bemerkungen. Deutsche Entomologische Zeitschrift 45: 154-155. Felsche, C. 1909. Einige coprophage Scarabaeiden, welche die Herren Oscar Neumann und Baron von Erlangen auf ihrer Reise in Abyssinien gesammelt haben. Deutsche Entomologische Zeitschrift 53: 448-451. Flach, K. 1879. Trox haroldi n. sp. Deutsche Entomologische Zeitschrift 23: 155-156. Gené, J. 1836. De quibusdam insectis Sardiniae novis aut minus cognitis. Memoralia Regiae Scientiarum Academiae Taurinensis 39: 161-199. Geoffroy, E. L. 1762. Histoire abrégée des insectes qui se trouvent aux environs de Paris, dans laquelle ces animaux sont rangés suivant un ordre métodique. Tome première. Durand; Paris. 523 p. Germar, E. F. 1824. Insectorum species novae aut minus cognitae, descriptionibus illustratae. Impensis J. C. Hendelii et Filii; Halle. 624 p. Germar, E. F. 1848. Beiträge zur Insectenfauna von Adelaide. Linnaea Entomologica 3: 153-247. Gerstaecker, C. E. A. 1866. Beschreibung neue Coleoptera. Archiv für Naturgeschichte 33(1): 44. Gerstaecker, C. E. A. 1873. Gliederthier-Fauna des Sansibar-Gebietes (Insekten, Arachniden, Myriapoden und Isopoden). In: Baron Carl Claus von der Decken’s Reisen in Ost-Afrika in den Jahren 1859-1865. Bd. 3, Abt. 2. Wissenschaftliche Ergebnisse, Abt. 2. C. F. Winter; Leipzig und Heidelberg. 542 p. CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 25 Gmelin, J. F. 1790. Caroli a Linné, systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima tertia, aucta, reformata. Tom. I, Pars IV, Classis V. Insecta. G. E. Beer; Lipsiae. p. 1517-2224. Goeze, J. A. E. 1777. Entomologische Beiträge zu des Ritter Linné zwölften Ausgabe des Natursystems. Erster Teil. Wiedemann; Leipzig. 736 p. Gutiérrez, A. R. 1950. Scarabaeidae del norte de Chile. Anales de la Sociedad Científica Argentina 149: 52-75. Haaf, E. 1953. Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). Entomologische Arbeiten aus dem Museum Georg Frey 4: 309-346. Haaf, E. 1954a. Die afrikanischen und orientalischen Arten der Gattung Trox (Col. Scarab.). Entomologische Arbeiten aus dem Museum Georg Frey 5(1): 326-393. Haaf, E. 1954b. Die australischen Arten der Gattung Trox (Col. Scarab.). Entomologische Arbeiten aus dem Museum Georg Frey 5: 691-740. Haaf, E. 1954c. Eine neue Trox-Art aus Süd-Afrika. Durban Museum Novitates 4(8): 97-99. Haaf, E. 1955a. Troginae (Coleoptera, Lamellicornia). Parc National de l’Upemba. I. Mission G. F. de Witte 1946-1949. Institut des Parcs Nationaux du Congo Belge 38(3): 45-50. Haaf, E. 1955b. Contributions à l’étude de la faune entomologique du Ruanda-Urundi (Mission P. Basilewsky 1953). Annales du Muséum Congo Belge (8), Zoologie 36: 87. Haaf, E. 1957a. Drei neue Trox-Arten (Col. Scarab.). Entomologische Arbeiten aus dem Museum Georg Frey 8: 692-695. Haaf, E. 1957b. On new Australian species of Trox. Memoirs of the National Museum of Victoria 21: 117-120. Haaf, E. 1958a. Coleoptera: Trogidae. In: R. Hanström, P. Brinck, and G. Rudebeck (eds.), South African Animal Life: Results of the Lund University expedition in 1950-1951, vol. 4. Statens Naturvetenskapliga Forskningsråd; Lund. 508 p. Haaf, E. 1958b. Nachtrag zur Revision der australischen Arten der Gattung Trox (Col. Scarab.). Entomologische Arbeiten aus dem Museum Georg Frey 9: 1078-1082. Haaf, E. 1959. Troginae und Sisyphini. In: Parc National de la Garamba. Mission H. de Saeger 1949- 1952. Institut des Parcs Nationaux du Congo Belge 15(2): 41-47. Haaf, E. 1960. XIV. Coleoptera Scarabaeidae: Trox et Sisyphus. In: Mission zoologique de l’I.R.S.A.C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). Annales du Muséum Congo Belge (8), Zoologie 81: 242-244. Harold, E. 1868. Diagnosen neuer Coprophagen. Coleopterologische Hefte 4: 79-86. Harold, E. 1871a. Beschreibung neue Coleoptera. Diagnosen neuer Coprophagen. Coleopterologische Hefte 7: 110-117. Harold, E. 1871b. Verzeichniss der von Dr. Beccari in Bogas gesammelten coprophagen Lamellicornien. Coleopterologische Hefte 8: 1-28. Harold, E. 1871c. Diagnosen neuer Coprophagen. Coleopterologische Hefte 8: 114-116. Harold, E. 1872. Monographie der Gattung Trox. Coleopterologische Hefte 9: 1-192. Harold, E. 1873. [Trox graecus Mulsant & Rey = T. transversus Reiche]. Coleopterologische Hefte 11: 118. Harold, E. 1880. Beschreibung neuer, auf seiner von der Akademie unterstützen Reise in Ost-Afrika; gesammelter Coleopteren. Monatsberichte der Akademie der Wissenschaften zu Berlin; p. 260- 270. Heller, K. M. 1914. Neue papuanische Käfer. Deutsche Entomologische Zeitschrift 58: 305-318. Herbst, J. F. W. 1783. Kritisches Verzeichniß meiner Insektensammlung. Archiv der Insectengeschichte, Bd. 4: 1-72, 1-23 tab. Herbst, J. F. W. 1790. Natursystem aller bekannten in- und ausländischen Insekten, als eine Forsetzung der von Büffonschen Naturgeschichte. Der Käfer. Theil 3. Joachim Pauli; Berlin. 325 p. Horn, G. H. 1874. Revision of the species of Trox of the United States. Transactions of the American Entomological Society 5: 1-12. Howden, H. F., and P. Vaurie. 1957. Two new species of Trox from Florida. American Museum Novitates 1818: 1-6. 26 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK ICZN (International Commission on Zoological Nomenclature) 1999. International Code of Zoological Nomenclature, Fourth Edition. The International Trust for Zoological Nomenclature; London. 306 p. Illiger, J. C. W. 1802a. Nachtrag und Berichtigungen zu dem Verzeichnisse der Käfer Preussens. Magazin für Insectenkunde 1[1801]: 1-94. Illiger, J. C. W. 1802b. Olivier’s Entomologie oder Naturgeschichte der Insekten mit ihren Gattungsund Art Merkmalen, ihren Beschreibung und Synonymie. Käfer. Uebersetzt mit Anmerkungen durchgängig begleitet. Zweiter Theil. K. Reichard; Braunschweig. 266 p. Kawai, S. 2006. A new species of the genus Omorgus Erichson, 1847 (Coleoptera, Trogidae) from South Africa. Kogane no. 7: 1-4. Kawai, S. 2006. Rare Scarabaeoidea of the world [3]. Omorgus dohrni (Harold, 1872) (Coleoptera, Trogidae). Saikaku-Tsûshin no. 13: back cover (English title, text in Japanese). Kawai, S. 2009. A new species of the genus Omorgus Erichson, 1847 (Coleoptera, Trogidae) from Australia. Kogane no. 10: 73-76. Klug, J. F. C. 1855. Forsetzung der Diagnosen der neuen (und bereits seit mehreren Monaten vollständig gedruckten) Coleopteren, welche die Insectensendungen des Hrn. Dr. Peters von Mossambique enthalten hatten, von der Familie der Staphylinii an bis den Lamellicornia, diese mit eingeschlossen. Berichten der Akademie der Wissenschaften (Berlin) 20: 643-660. Kolbe, H. J. 1891. Aufzahlung den von Herr Dr. Hans Meyer in Jahre 1889 in Gebiete das Kilimandscharo- und Ugueno-Gebirges gesammelten Coleopteren. Stettiner Entomologische Zeitung 52: 18-38. Kolbe, H. J. 1904. Über einige interessante Lamellicornier und Tenebrioniden Afrikas. Berliner Entomologische Zeitschrift 49: 282-302. Kolbe, H. J. 1908. Dynastidae, Cetoniidae und Scarabaeidae. In: L. Schultze, Zoologische und anthropologische Ergebnisse einer Forschungsreise im westlichen und zentralen Südafrika. Erster Band. Denkschriften des Medizinisch-Naturwissenschaftlichen Gesellschaft (Jena) 13: 121-132. Kolbe, H. J. 1914. Scarabaeiden. In: H. Schubotz. Wissenschaftliche Ergebnisse der deutsche in dem Zentral Afrika Expeditionen, Bd. 5, Lieferung 3. Klinkhardt und Biermann; Leipzig. p. 261- 416. Král, D., and V. Kubán. 2012. Afromorgus reiterorum sp. nov. (Coleoptera: Trogidae) from Socotra Island. Acta Entomologica Musei Nationalis Pragae 52 (Supplementum 2): 147-152. Krell, F.-T. 2006. Fossil record and evolution of Scarabaeoidea (Coleoptera: Polyphaga). p. 120-143. In: M. L. Jameson, and B. C. Ratcliffe. (eds.). Scarabaeoidea in the 21st Century: A Festschrift Honoring Henry F. Howden. Coleopterists Society Monograph no. 5: 216 p. Krell, F.-T. 2007. Catalogue of fossil Scarabaeoidea (Coleoptera: Polyphaga) of the Mesozoic and Tertiary – Version 2007. Denver Museum of Nature & Science, Technical Report 2007-8: 1-79. Krynicki, J. 1832. Enumeratio coleopterorum Rossiae meridionalis et praecique in Universitatis Caesareae Charkoviensis circulo obvenientium, quae annorum 1827-1831 spatio observavit, P. P. E. Ioannes Krynicki. Bulletin de la Société Impériale des Naturalistes de Moscou 5: 65-179. Lacordaire, J. P. 1856. Histoire naturelle des insectes. Genera des Coléoptères, ou exposé méthodique et critique de tous les genres proposes jusqu’ici dans cet ordre d’insectes. Tome troisième contenant les familles des pectinicornes et lamellicornes. Librairie Encyclopédique de Robet; Paris. 594 p. Laicharting, J. N. E. 1781. Verzeichniss und Beschreibung der Tyrolen-Insecten, I. Theil. Käferartige Insecten. I. Band. Füessly; Zurich. 248 p. Lansberge, J. W. 1886. Scarabaeides, Buprestides et Cerambycides de l’Afrique occidentale envoyés ou Musée Leyde par MM. Veth et Van der Kellen. Notes from the Leyden Museum 8: 69-120. Lansberge, J. W. 1887. Trogides nouveaux. Notes from the Leyden Museum 9: 199-211. Latreille, P. A. 1810. Considerations générales sur l’ordre naturel des animaux composant les classes des crustacés, des arachnides et des insectes; avec un tableau méthodique de leur genres, disposés en familles. F. Schoell; Paris. 444 p. LeConte, J. L. 1854. Descriptions of the species of Trox and Omorgus inhabiting the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 7: 211-216. Lewis, G. 1895. On the lamellicorn Coleoptera of Japan, and notices of others. Annals and Magazine of Natural History (6) 16: 374-408. CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 27 Linné, C. 1758. Systema Naturae per Regna tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Impensis Direct. Laurentii Salvii; Stockholm. 824 p. Linné, C. 1767. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio duodecima, reformata. Tomus I, Pars II. Laurentii Salvii; Stockholm. p. 533-1327. Loomis, H. F. 1922. New species of the coleopterous genus Trox. Journal of the Washington Academy of Sciences 12: 132-136. MacLeay, W. J. 1864. Descriptions of new genera and species of Coleoptera from Port Denison. Transactions of the Entomological Society of New South Wales 1: 106-130. MacLeay, W. J. 1871. Notes on a collection of insects from Gayndah. Transactions of the Entomological Society of New South Wales 2: 79-205. MacLeay, W. J. 1888. The insects of King’s Sound and its vicinity, 2. The lamellicorns. Proceedings of the Linnean Society of New South Wales (2) 3: 897-924. MacLeay, W. S. 1819. Horae entomologicae; or essays on the annulose animals. Containing general observations on the geography, manners, and natural affinities of the Insects which compose the genus Scarabaeus of Linnaeus; to which are added a few incidental remarks on the genera Lucanus and Hister of the same author. With an appendix and plates. Vol. 1, Pt. 1. S. Bagster; London. 160 p. MacLeay, W. S. 1826. Appendix: Annulosa, Mandibulata. p. 2-33. In: P. King (ed.), Narrative of a survey of the intertropical and western coasts of Australia performed between the years 1818 and 1822 by Captain Phillip P. King, R.N., F.R.S., F.L.S., and member of the Royal Asiatic Society of London, with an appendix relating to hydrography and natural history. Vol. II. John Murray; London [larger version in 1827]. Marchand, E. 1902. Description de deux arthropodes nouveaux provenant du Soudan francais. Bulletin de la Société de l’Ouest de France 2: 331-343. Marsham, T. 1802. Entomologia Britannica, sistens insecta Britanniae indigena, secundum methodum Linnaeanam disposita. Tomus I. Coleoptera. Wilks and Taylor; London. 547 p. Masumoto, K. 1996. Coprophagid-beetles from Northwest Thailand (XI) (Coleoptera, Trogidae). The Entomological Review of Japan 51(1): 9-13. Masumoto, K., and M. Kiuchi. 1995. A new Trox species (Coleoptera, Trogidae) from Amamioshima of the Nansei Islands, Southwest Japan. Elytra 23(2): 205-208. Masumoto, K., T. Ochi, and C.-L. Li. 2005. A revision of the Taiwanese species of the family Trogidae (Coleoptera, Scarabaeoidea). Elytra 33(1): 229-243. Mawdsley, J. R. 1993. The entomological collection of Thomas Say. Psyche 100: 163-171. Melsheimer, F. E. 1846. Descriptions of new species of Coleoptera of the United Strates. Proceedings of the Academy of Natural Sciences of Philadelphia 2: 134-160. Mikšiæ, R. 1958. Eine neue Trox-Art aus dem Togoland. Entomologische Berichten 18: 15-16. Motschulsky, V. 1860. Diagnose d’insectes nouveaux des rives du fl. Amur et de la Daourie méridionale. Etudes Entomologiques 9: 39-41. Mulsant, E., and C. Rey. 1871. Histoire naturelle des Coléoptères de France. Tribu des Lamellicornes. Annales de la Société d’Agriculture, Histoire Naturelle et Arts Utiles de Lyon (4) 2[1869]: 241- 650; 3[1870]: 155-480. Mutchler, A. J. 1925. Coleoptera from the Williams Galapagos expeditions. Zoologica 5(20): 219-240. Nakane, T. 1954. New or little known Coleoptera from Japan and its adjacent regions. X. Descriptions of some new species from Shikoku, Japan. Transactions of the Shikoku Entomological Society 4: 7-15. Nakane, T. 1983. New or little-known Coleoptera from Japan and its adjacent regions. 36. Fragmenta Coleopterologica 35/37: 139-150. Nakane, T., and K. Tsukamoto. 1955. [On the family Trogidae in Japan. Studies on the Lamellicorn Coleoptera II]. Transactions of the Kyoto Entomological Society 4: 85-92 (in Japanese, English abstract). Nikolaev, G. V. 2000. Novoe podsemeistvo plastinchatousykh zhukov (Coleoptera, Scarabaeidae) iz nizhnego mela Zabaikalia i Mongolii i ego polozhenie v sisteme nadsemeistva. Paleontologicheskii Zhurnal (Moscow) 2000(4): 63-66 (in Russian). 28 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK Nikolaev, G. V. 2005. Omorgini (Coleoptera, Scarabaeinae, Troginae) – novaia triba plastinchatousykh zhukov. Evraziiskii Entomologicheskii Zhurnal [Euroasian Entomological Journal] 4(4): 321-322 (in Russian). Nikolaev, G. V. 2007. Mezozoiskii etap evoliucii plastinchatousykh (Insecta: Coleoptera: Scarabaeoidea). Kazak Universiteti; Almaty. 222 p. Nomura, S. 1937. On the genus Trox of Japan (Scarabaeidae) with description of a new species. Nippon no Kôchû 1: 77-86 (in Japanese, English abstract). Nomura, S. 1961. Some new species of the Coleoptera from Japan. Tôhô-Gakuhô 11: 70-89. Nomura, S. 1973. Notes on the coprophagous Lamellicornia from Taiwan. Entomological Review of Japan 25(1-2): 37-52. Ochi, T., and M. Kawahara. 2002. Taxonomic notes on some Japanese coprophagous lamellicorn beetles (Coleoptera, Scarabaeoidea). IV. A new subspecies of Trox setifer and a new record of T. opacotuberculatus from Tsushima Is. Kogane no. 3: 53-58. Ochi, T., M. Kawahara, and M. Inagaki. 2008. A new species of the genus Trox from Yakushima Is., southwest Japan (Coleoptera, Scarabaeoidea, Trogidae). Kogane no. 9: 135-138. Ochi, T., M. Kon, and M. Bai. 2010. A new species of the genus Trox from Sichuan, China (Coleoptera: Scarabaeoidea: Trogidae). Entomological Review of Japan 65(1): 131-134. Ochi, T., M. Kon, and M. Kawahara. 2005. Trogidae (Coleoptera) from Borneo, with description of a new species of Trox and a new record of T. scaber. Kogane no. 6: 65-70. Olivier, A. G. 1789. Entomologie ou Histoire naturelle des insectes, avec leurs caractères génériques et spécifiques, leur description, leur synonymie et leur figures enluminées. Coléoptères. Tome premier. Baudoin; Paris. 497 p. [genera paginated separately]. Pallas, P. S. 1781-1806. Icones Insectorum praesertim Russiae Sibiriaeque peculiarium quae collegit et Descriptionibus illustravit. Wolfgangi Waltheri; Erlangae. 104 p. [issued in parts]. Paulian, R. 1933. Coprophages asiatiques nouveaux ou peu connus (Col.). Bulletin de la Société Entomologique de France 38: 205-208. Paulian, R. 1937. Faune entomologique de Madagascar. Bulletin de la Académie de Malgache (n. s.) 19[1936]: 129-143. Paulian, R. 1948. Coleoptera, Scarabaeidae: Trogidae, Geotrupinae, Dynamopinae, Hybosorinae, Coprinae, Aphodiinae. In: British Museum (Natural History) Expeditions to south-west Arabia 1937-1938. British Museum (Natural History); London. p. 141-155. Paulian, R. 1981. Faune de Madagascar, 56. Insectes Coléoptères: Trogidae et Hybosoridae. CNRS; Paris. 28 p. Paulus, H. F. 1978. Trox dhaulagiri n. sp. aus Nepal. Senckenbergiana Biologica 53(1-2): 55-58. Paykull, G. 1798. Fauna Suecica. Insecta. Tomus I. J. E. Edman; Upsaliae. 358 p. Péringuey, L. 1901-1903. Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society 12: 1-920 [1901: 1-563; 1902: 564-896; 1903: 897-920]. Péringuey, L. 1908. Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Additions and Corrections. Transactions of the South African Philosophical Society 13: 547-752. Petrovitz, P. 1975. Zwei neue Trox-Arten. Opuscula Zoologica 132: 1-3. Petrovitz, P. 1980. Österreichische entomologische Expeditionen nach Persien und Afghanistan. Beiträge zur Coleoptera Fauna, 12. Weiteres über Lamellicornia aus Iran. Annalen des Naturhistorischen Museums in Wien 83: 597-638. Pittino, R. 1983. Trox (s.str.) granulipennis Fairm. and allied “taxa”, with two new species from Mediterranean area. Giornale Italiano di Entomologia 1: 265-284. Pittino, R. 1985. Four new Oriental species of the genus Trox (s.str.) F. Giornale Italiano di Entomologia 2(10): 327-340. Pittino, R. 1987. New Coleoptera Trogidae from South America. Giornale Italiano di Entomologia 3: 377-397. Pittino, R. 1991. On some Palaearctic “taxa” allied to Trox hispidus (Pontoppidan) with a brachypterous new species from Italy, Malta, Crete and the Balkan peninsula (Coleoptera, Trogidae). Bollettino dell’ Associazione Romana di Entomologia 45[1990]: 57-87. CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 29 Pittino, R. 2005. Four new Asian species of Omorgus Erichson, 1847 (Coleoptera, Scarabaeoidea, Trogidae). Kogane no. 6: 71-79. Pittino, R. 2006. Trogidae (p. 26-28, 79-81). In: I. Löbl, and A. Smetana. (eds.), Catalogue of Palaearctic Coleoptera, Vol. 3. Apollo Books; Stenstrup. 690 p. Pittino, R. 2010. A new genus and species of Trogidae MacLeay, 1819 from Madagascar, and a new species of Omorgus Erichson, 1847 from Argentina (Coleoptera, Trogidae). Kogane no. 11: 75-83. Pittino, R., and S. Kawai. 2006. Notes on the genus Trox Fabricius, 1775 from East Asia. I. Description of the males of Trox (Trox) cambodjanus and T. (T.) poringensis and definition of the “terrestris” species group (Coleoptera, Trogidae). Kogane no. 7: 5-10. Pittino, R., and S. Kawai. 2010. Redescription of Omorgus marshalli (Haaf, 1957) (Coleoptera, Trogidae) from Australia. Kogane no. 11: 85-88. Pontoppidan, E. 1763. Den Danske atlas eller Konge-Riget Danemark, med dets naturlige edenskaber, elementer, inbyggere, vaexter, dyr og affodninger, dets gamle tildraglser og naervaerende omstaendingheder I aller provinter, staeder, kirkzer, slotte og herre-gaarde. Tomus I. A. H. Godiche; København. 723 p. Preudhomme de Borre, F. P. C. A. 1886. Catalogue des Trogides décrits jusque’ à ce jour précédé d’ un synopsis de leurs genres et d’ une esquisse de leur distribution géographique. Annales de la Société Entomologique de Belgique 30: 56-82. Ratcliffe, B. C. 1978. A new species of Trox from the Amazon basin with new distributional records for Central American Trox. Acta Amazonica 8(2): 299-302. Reiche, L. J. 1853. Note sur l’Hybosorius orator et sur le Trox granulatus, et description d’une nouvelle espèce. Annales de la Société Entomologique de France (3) 1: 87-90. Reiche, L. J. 1856. In: L. J. Reiche, and F. Saulcy, Espèces nouvelles ou peu connues de Coléoptères, recueillis par F. de Saulcy, membre de l’Institut, dans son voyage en Orient, et décrites par MM. Reiche et Félicien de Saulcy (Suite). Annales de la Société Entomologique de France (3) 4: 353- 422. Reiche, L. J. 1861. Coléoptères nouveaux recueillis en Corse par M. E. Bellier de la Chavignerie. Annales de la Société Entomologique de France (4) 1: 201-210. Reitter, E. 1892. Bestimmungs-Tabellen der Lucaniden und coprophagen Lamellicornen der palaearktischen Faunengebietes. Verhandlungen des Naturforschenden Vereins in Brünn 30[1891]: 141-262. Reitter, E. 1904. Sechzehn neue Coleopteren aus Europa und den angenzenden Ländern. Wiener Entomologische Zeitung 23: 150-160. Ren, G.-D. 2003. Taxonomic studies of the family Trogidae (Coleoptera: Scarabaeoidea) from China. Entomotaxonomia 25: 109-117 (in Chinese, English abstract). Ritcher, P. O. 1966. White grubs and their allies. A study of North American scarabaeid larvae. Oregon State University Press; Corvallis. 219 p. Robinson, M. 1940. Studies in the Scarabaeidae, 2. Transactions of the American Entomological Society 66: 141-159, pl. 4. Robinson, M. 1941a. Studies in the Scarabaeidae of North America, 1, 2. Transactions of the American Entomological Society 67: 127-136. Robinson, M. 1941b. A new species of Trox from Texas. Entomological News 52: 134-135. Robinson, M. 1941c. Notes on some rare Scarabaeidae with the description of one new species. Entomological News 52: 227-232. Robinson, M. 1946. Studies in the Scarabaeidae, 3. Transactions of the American Entomological Society 72: 49-59. Robinson, M. 1948. A new genus of Scarabaeidae. Notulae Naturae no. 211: 1-2. Rossi, P. 1790. Fauna Etrusca sistens insecta quae in provinciis Florentina et Pisana praesertim collegit Petrus Rossius in regio pisano athenaeo. Tomus primus. Typis Thomae Masi et Sociorum; Liburni. 272 p. Rossi, P. 1792. Mantissa Insectorum, exhibens species nuper in Etruria collectas, adiectis Faunae Etruscae illustrationibus, ac emendationibus. Tom. I. Typografia Polloni; Pisis. 148 p. Roth, J. R. 1851. Diagnosen neuer Coleoptera aus Abyssinien. Archiv für Naturgeschichte 17(1): 115-133. 30 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK Say, T. 1823. Descriptions of coleopterous insects collected in the late expedition to the Rocky Mountains; performed by order of Mr. Calhoun, Secretary of War, under the command of Major Long. Journal of the Academy of Natural Sciences of Philadelphia 3(1): 238-282. Say, T. 1824. Descriptions of Coleoptera. In: W. H. Keating, Narrative of an expedition to the source of St. Peters’ River, & c., under the command of Stephen H. Long, Major U.S.T.E., Vol. 2. Philadelphia. p. 268-378. Say, T. 1825. Descriptions of new species of coleopterous insects inhabiting the United States. Journal of the Academy of Natural Sciences of Philadelphia 5(1): 160-204. Say, T. 1831. New species of North American insects found by Joseph Barabino chiefly in Lousiana. New Harmony, Indiana; 16 p. Say, T. 1835. Descriptions of North American coleopterous insects and observations on some already described. Boston Journal of Natural History 1: 151-203. Schmidt, G. 1936. Studien über Trox hispidus Pontopp. und seine Formen. Entomologische Blätter 32: 75-78. Scholtz, C. H. 1978. The occurrence and distribution of Trox in the Kruger National Park. Koedae 21: 81-89. Scholtz, C. H. 1979a. Taxonomic revision and distribution of the genus Trox in South West Africa. Cimbebasia (A) 5(3): 179-200. Scholtz, C. H. 1979b. The “horridus” group of Trox F. species in South Africa. Journal of the Entomological Society of Southern Africa 42(2): 169-180. Scholtz, C. H. 1980a. Monograph of the genus Trox F. of subsaharan Africa. Cimbebasia, Memoir 4: 1-104. Scholtz, C. H. 1980b. Trogidae. In: W. Wittmer (ed.). Fauna of Saudi Arabia, Vol. 2. Basel. p. 137- 140. Scholtz, C. H. 1981. Aptery in Trox: morphological changes and their relationship to habitat. Journal of the Entomological Society of Southern Africa 44(1): 83-87. Scholtz, C. H. 1982. Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Department of Agriculture and Fisheries, Republic of South Africa, Entomology Memoir no. 54: 1-27. Scholtz, C. H. 1983. A new species of Trox F. from Senegal with new distributional records for other African species. Journal of the Entomological Society of Southern Africa 46: 206-208. Scholtz, C. H. 1986a. Phylogeny and systematics of the Trogidae. Systematic Entomology 11: 355- 363. Scholtz, C. H. 1986b. Revision of the genus Trox Fabricius of the Australasian region. Australian Journal of Zoology, Supplement Series 125: 1-99. Scholtz, C. H. 1986c. A new species of Trox Fabricius from Zambia. The Coleopterists Bulletin 40(1): 29-31. Scholtz, C. H. 1990a. Phylogenetic trends in the Scarabaeoidea. Journal of Natural History 24: 1027-1066. Scholtz, C. H. 1990b. Revision of the Trogidae of South America. Journal of Natural History 24: 1391-1456. Scholtz, C. H. 1991a. Descriptions of larvae of Australian Omorgus Erichson, with implications for the phylogeny of the Trogidae. Invertebrate Taxonomy 5: 827-835. Scholtz, C. H. 1991b. A new species of Omorgus Erichson from West Africa. The Coleopterists Bulletin 45(2): 112-114. Scholtz, C. H. 1993a. Descriptions of South American Trogidae larvae. The Coleopterists Bulletin 47(2): 209-214. Scholtz, C. H. 1993b. Descriptions of larvae of African Trogidae, with implications for the phylogeny of the family. African Entomology 1: 1-13. Scholtz, C. H., and J.-P. Lumaret. 1991. Descriptions of European Trox larvae. The Coleopterists Bulletin 45(4): 317-322. Scholtz, C. H., and S. Peck. 1990. Description of a Polynoncus Burmeister larva, with implications for phylogeny of the Trogidae. Systematic Entomology 15: 383-389. Scholtz, C. H., D. J. Browne, and J. Kukalová-Peck. 1994. Glaresidae, archaeopteryx of the Scarabaeoidea (Coleoptera). Systematic Entomology 19: 259-277. CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 31 Scholtz, C. H., D. d’Hotman, and A. Nel. 1987a. Glaresidae, a new family of Scarabaeoidea (Coleoptera) to accommodate the genus Glaresis Erichson. Systematic Entomology 12: 345-354. Scholtz, C. H., D. d’Hotman, and A. Nel. 1987b. Systematic position of the South American genus Cryptogenius Westwood. The Coleopterists Bulletin 41(2): 193-199. Scholtz, C. H., D. J. G. Inward, and M. D. Kerley. 2007. Description of a new Palaearctic Trox Fabricius species (Coleoptera: Trogidae) from Thailand, with a review of the Trox opacotuberculatus species-group and its biogeography. Insect Systematics and Evolution 38: 93- 103. Schultze, W. 1915. Beitrag zur coleopteren Fauna der Philippinen. Philippine Journal of Science (D) 10: 271-279. Scudder, S. H. 1879. The fossil insects collected in 1877 by Mr. G. M. Dawson in the interior of British Columbia. In: G. M. Dawson, Preliminary report on the physical and geological features of the southern portion of the interior of British Columbia. Geological Survey of Canada, Report of Progress 1877-78: 175B-185B. Smith, A. B. T., D. C. Hawks, and J. M. Heraty. 2006. An overview of the classification and evolution of the major scarab beetle clades (Coleoptera: Scarabaeoidea) based on preliminary molecular analyses (p. 35-46). In: M. L. Jameson, and B. C. Ratcliffe. (eds.), Scarabaeoidea in the 21st Century: A Festschrift Honoring Henry F. Howden. Coleopterists Society Monograph no. 5: 216 p. Steiner, W. E. 1981. A new species of Trox from Peru, with notes on the biology and distribution of Trox aricensis Gutierrez. The Coleopterists Bulletin 35(2): 205-210. Struempher, W. P., and C. H. Scholtz. 2011. Description of a new, possibly extinct, species of flightless Trox F. (Coleoptera: Trogidae) from South Africa. Insect Systematics and Evolution 42(4): 337-348. Thunberg, C. P. 1787. Museum Naturalium Academiae Upsalensis. Cujus partem quartam. J. Edman; Upsaliae. 58 p. Van Dyke, E. C. 1953. The Coleoptera of the Galapagos Islands. California Academy of Sciences, Occasional Papers no. 22: 181 p. Vaurie, P. 1955. A revision of the genus Trox in North America. Bulletin of the American Museum of Natural History 106(1): 1-89. Vaurie, P. 1962. A revision of the genus Trox in South America. Bulletin of the American Museum of Natural History 124(4): 105-167. Walker, F. 1858. Characters of some apparently undescribed Ceylon insects. Annals and Magazine of Natural History (3) 2: 202-209. Waterhouse, C. O. 1875. The lamellicorn Coleoptera at present known to occur in Japan. Transactions of the Royal Entomological Society of London 1875: 71-116. Wickham, H. F. 1909. New fossil Coleoptera from Florissant. American Journal of Science (4) 28: 126-130. Wiedemann, C. R. W. 1821. [New taxa]. In: C. R. W. Wiedemann, and E. F. Germar, Neue exotische Käfer. Magazin der Entomologie 4: 107-183. Wiedemann, C. R. W. 1823. Zweihundert neue Käfer von Java, Bengalen und dem Vorgebirge der gutten Hoffnung. Zoologisches Magazin 2(1): 1-133. Wollaston, T. V. 1864. Catalogue of the coleopterous insects of the Canaries in the collection of the British Museum. Taylor and Francis; London. 648 p. Wollaston, T. V. 1867. Coleoptera Hesperidium, being an enumeration of the Coleopterous Insects of the Cape Verde Archipelago. John van Voorst; London. 285 p. Wollaston, T. V. 1877. Coleoptera Sanctae-Helenae. John van Voorst; London. 256 p. Zhang, Y.-W. 1988. Scarabaeidae. p. 227-248. In: F.-S. Huang, P.-Y. Wang, W.-Y. Yin, P.-Y. Yu, T.-S. Lee, C.-K. Yang, and X.-J. Wang (eds.), Insects of Mt. Namjagbarwa Region of Xizang. Science Press; Beijing. 621 p. (in Chinese, English abstract). Zoubkoff [= Zubkov], V. P. 1829. Notice sur un nouveau genre quelques nouvelles espèces de Coléoptères. Bulletin de la Société Impériale des Naturalistes de Moscou 1: 147-168. 32 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK APPENDIX: Supplemental literature not cited, but of importance to the study of Trogidae. Aston, P., and S. Kawai. 2010. Afromorgus pauliani (Haaf 1954), a second species of Trogidae (Coleoptera: Scarabaeoidea) recorded in Hong Kong. Hong Kong Entomological Society 2(1): 43. Bacchus, M. E. 1978. A catalogue of the type-specimens of the Scarabaeinae (Scarabaeidae) and the smaller lamellicorn families (Coleoptera) described by G. J. Arrow. Bulletin of the British Museum (Natural History), Entomology series 37(3): 97-115. Báguena, L. 1960. Los Troginae ibéricos. Graellsia 18: 147-151. Baraud, J. 1985. Coléoptères Scarabaeoidea. Faune du Nord de l’Afrique du Maroc au Sinaï. Encyclopédie Entomologique 46. P. Lechevalier; Paris. 652 p. Baraud, J. 1992. Coléoptères Scarabaeoidea d’Europe. Fauna de France 78. Fédération française des Sociétés de Sciences Naturelles et Société Linnéenne de Lyon. 856 p. Bates, H. W. 1886-1890. Biologia Centrali-Americana. Insecta. Coleoptera, Vol. 2, Pt. 2. Pectinicornia and Lamellicornia. Taylor and Francis; London. 423 p. [Trogidae in 1887: 116-118 + suppl. vol. 2, pt. 2; 1889: 395]. Bedel, L. 1881. Faune des Coléoptères du bassin de la Seine. Annales de la Société Entomologique de France, Hors séries; 360 p. Bedel, L. 1884. Recherches sur la synonymie des Coléoptères de l’ancien monde. Abeille 28: 150-156. Berg, F. G. C. 1898. Sobre los enemigos pequeños de la langosta peregrina Schistocerca paranensis (Burm.). Communicaciones del Museo Nacional de Buenos Aires 1: 25-30. Bezdek, A., and J. Hájek. 2009. Catalogue of type specimens of beetles (Coleoptera) deposited in the National Museum, Prague, Czech Republic. Scarabaeoidea: Bolboceratidae, Geotrupidae, Glaphyridae, Hybosoridae, Ochodaeidae and Trogidae. Acta Entomologica Musei Nationalis Pragae 49(1): 297-332. Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America, pt. 2. Scarabaeidae, Troginae. Bulletin of the U. S. National Museum 185: 218-219. Blackwelder, R. E., and R. M. Blackwelder. 1948. Fifth supplement to the Leng Catalogue of Coleoptera of America, north of Mexico. John D. Sherman; Mt. Vernon, New York. 87 p. Blatchley, W. S. 1910. The Coleoptera or beetles of Indiana. The Indiana Department of Geology and Natural Resources, Bulletin no. 1: 1-1386. Blatchley, W. S. 1928. The Scarabaeidae of Florida. Florida Entomologist 12: 63-65. Britton, E. B. 1970. Coleoptera. In: D. F. Waterhouse (ed.). The insects of Australia. Melbourne University Press; p. 495-621. Brivio, P. C. 1959. Contributo alla conoscenza della fauna coleotterologica della Guinea portuguese, 3. Atti della Società Italiana di Scienze Naturali 98: 372-378. Bruch, C. 1911. Catalogo systemática de los coleópteros de la República Argentina. Revista del Museo de La Plata 17: 143-260. Burakowski, B., M. Mroczkowski, and J. Stefanska. 1983. Chraszce. Coleoptera (Scarabaeoidea etc.). Katalog Fauny Polski, cz. 23, r. 9. PWN; Warszawa. 294 p. Campbell, J. M. 1991. Troginae. p. 150-151. In: Y. Bousquet (ed.), Checklist of beetles of Canada and Alaska. Agriculture Canada Publication 1861/E: 430 p. Candèze, E. C. A. 1870 (1871). [Trox in wool from Argentina]. Annales de la Société Entomologique de Balgique, Comptes-Rendus des Séances 14: xxiii. Cassis, G., and T. A. Weir. 1992. Trogidae. p. 30-40. In: W. W. K. Houston (ed.), Zoological Catalogue of Australia, Vol. 9. Coleoptera: Scarabaeoidea. AGPS; Canberra. 544 p. Caveney, S., and C. H. Scholtz. 1993. Evolution of ommatidium structure in the Trogidae. Systematic Entomology 18: 1-10. Chalumeau, F., and L. Gruner. 1974. Scarabaeoidea des Antilles francaises. Annales de la Société Entomologique de France (n. s.) 10(4): 781-819. Conil, P. A. 1880. Etudes sur l’Acridium paranense Burm., ses variétés et plusieurs innetes qui le détruisent. Sociedad de Zoología Argentina 3: 177- 257. Costa, C., S. A. Vanin, and S. A. Casari-Chen. 1988. Larves de Coleoptera de Brasil. Museu de Zoologia da Universidade de São Paulo. 282 p. CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 33 Dawson, R. W. 1923. A synopsis of the Scarabaeidae of Nebraska. University of Nebraska Studies 22(3-4): 163-244. Decelle, J. 1972. La faune terrestre de l’Île de Sainte Héléne, 2. Insectes, 9. Coleoptera, 6. Fam. Trogidae. Annales de la Société Zoologique de Belgique 192: 114. Deloya, C. 2000. Distribucion de la familia Trogidae en Mexico. Acta Zoologica Mexicana (n. s.) 81: 63-76. Deloya, C., and L. Quiroz-Robledo. 1992. A new southern record of Trox aequalis Say 1831 in Mexico. The Coleopterists Bulletin 46(4): 420. Denier, P. 1936. Estado natural de mis conocimientos acerca del “champi” (Trox suberosus F.). Memoria de la Comisión Central de Investigaciones Sobre La Langosta (Argentina). Diéguez, V. M., and R. Gómez. 2004. Aporte al conocimiento de los Trogidae de la Argentina (Coleoptera). Revista de la Sociedad Entomológica Argentina 63(1-2): 92-95. Duftschmid, C. 1825. Fauna Austriae. Dritter Theil. Verlag der k. k. priv. akademischen Kunst-, Musik- und Buchhandlung; Linz. 289 p. Endrödi, S. 1956. Lamellicornia (Coleoptera 4). 4, fuzet. Fauna Hungariae 12. Akademiai Kiado; Budapest. 180 p. Endrödi, S. 1971. The scientific results of the Hungarian zoological expeditions to Tanganyika, 16. Coleoptera: Lamellicornia. Folia Entomologica Hungarica 24(26): 289-311. Endrödi, S. 1976. Lamellicornia (Col.) aus Ghana, 2. Annales Historico-Naturales Musei Nationalis Hungarici 68: 155-164. Endrödi, S. 1977. Neue südwestafrikanische Aphodiinen gesammelt von Dr. H. Roer. Bonner Zoologische Beiträge 28(1/2):155-159. Fourcroy, A. F. de 1785. Entomologia parisiensis, sive catalogus insectorum, quae in agro parisiensi reperiuntur. Tome 1. Paris. 231 p. Frey, G., and A. Villiers. 1971. Contribution à l’étude biologique du Sénégal septentrional. 10. Coléoptères Trogidae, Geotrupidae et Scarabaeidae Onthophagini. Bulletin de l’Institut Fondamental d’Afrique Noire, Série A, Sciences Naturelles (1) 33: 137-144. Gardner, J. C. M. 1946. A note on the larva of Trox procerus. Indian Journal of Entomology 8: 31- 32. Gebler, F. 1847. Verzeichniss der im Kolywano-Woskresenskischen Hüttenbezirke Süd-West Sibiriens beobachteten Kaefer mit Bemerkungen und Beschreibungen. Bulletin de la Société Impériale des Naturalistes de Moscou 20(2): 391-512. Gemminger, M., and E. Harold. 1869. Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. IV. E. H. Gummi; Monachii. p. 979-1346. Grimshaw, P. H. 1897. On some type specimens of Lepidoptera and Coleoptera in the Edinburgh Museum of Science and Art. Transactions of the Royal Society of Edinburgh 39: 1-11. Guérin de Méneville, F. E. 1844 (1829-1844). Iconographie du régne animal de G. Cuvier, ou représentation d’après nature de l’une des espèces les plus remarquables et souvent non encore figurées, de chaque genre d’animaux. Avec un texte descriptif mis au courant de la science. Ouvrage pouvant servir d’atlas à tous les traités de zoologie. III. Texte explicatif. Insectes. J. J. Baillière; Paris. 576 p. + 104 pls. Gyllenhal, L. 1808. Insecta Suecica descripta. Classis I. Coleoptera sive Eleuterata. Tomus I. F. J. Leverentz; Skara [Scaris]. 572 p. Harris, T. W. 1837. Characteristics of some previously described North American coleopterous insects in the collection of Mr. A. Halsey. Transactions of the Herts Natural History Society 1: 65-91. Hatch, M. H. 1971. The beetles of the Pacific Northwest, pt. 5. University of Washington Publications in Biology 16: 1-662. Hayward, K. J. 1936. Contribucion al conocimiento de la langosta Schistocerca paranensis Burm. y sus enemigas naturales. Memoria de la Comisión Central de Investigaciones Sobre La Langosta (Argentina). Heer, O. 1841. Fauna Coleopterorum Helvetica. Pars I. Impensis Orelii, Fuesslini et Sociorum; Turici [Zurich]. 652 p. [Fasc. III: 361-652 issued in 1841]. Heim de Balsac, H. 1952. Considerations sur une biocénose constituée au tour d’un nidde cigogne Ciconia ciconia en Lorraine. Alauda 20: 144-156. 34 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK Henriksen, K. L. 1925. Torbist larverne. Danmarks Fauna 29: 125-170. Henshaw, S. 1885. List of the Coleoptera of America, north of Mexico. American Entomological Society, Philadelphia. 161 p. Hoffmann, W. E. 1931. Coprophagous beetles from Hainan Island. Lignan Science Journal 10: 501- 502. Horion, A. 1958. Faunistik der mitteleuropäischen Käfer, 6. Lamellicornia. A. Feyel Verlag; Überlingen-Bodensee. 343 p. Horn, G. H. 1885. Descriptions of new North American Scarabaeidae. Transactions of the American Entomological Society 12: 117-118. Horn, G. H. 1887. Notes on Lachnosterna. Entomologica Americana 3: 141-145. Iablokoff-Khnzorian, S. M. 1967. Nasekomye zhestokrylye. Fauna Armianskoi SSR, Tom VI, Plastinchatousye (Scarabaeoidea). Akademia Nauk Armianskoi SSR; Yerevan. 223 p. Illiger, J. C. W. 1798. Verzeichniss der Käfer Preussens. Entworten von Johan Gottlieb Kugelann, Apotheker in Osterode. Mit einer Vorrede des Professor und Pagenhofmeisters Hellwig in Braunschweig, und dem angehängten Versuche einer natürlichen Ordnungs- und Gattungs-Folge der Insecten. J. J. Gebauer; Halle. 510 p. Illiger, J. C. W. 1803. Verzeichniss der in Portugall einheimischen Käfer. Erste Lieferung. Magazin für Insectenkunde 2[1802]: 186-258. Jacquelin du Val, P. N. C. 1859-1863. Genera des Coléoptères d’Europe comprenant leur classification en familles naturelles, la description de tous les genres, des Tableaux synoptiques destinés à faciliter l’étude, le Catalogue de toutes les espèces, de nombreux dessins au trait de caractères. Tome troisième. A. Deyrolle; Paris. 463 p. Jameson, M. L. 2002. Trogidae. p. 17-19. In: R. H. Arnett jr, M. C. Thomas, P. E. Skelley, and J. H. Frank. (eds.). American Beetles, Vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press; Boca Raton - London - New York - Washington DC. 861 p. Juøena, D. 1996. Contribution to the faunistics of Scarabaeoidea (Coleoptera) of Bohemia, Moravia and Slovakia. Klapalekiana 32: 27-32. Kawai, S. 2006. Rare Scarabaeoidea of the world [3]. Omorgus dohrni (Harold, 1872) (Coleoptera, Trogidae). Saikaku-Tsûshin no. 13: back cover (English title, text in Japanese). Kawai, S. 2007. First record of Polynoncus ecuadorensis (Vaurie, 1962) from Peru (Coleoptera, Trogidae). Kogane no. 8: 96. Kawai, S. 2009. First record of Afromorgus satorui (Kawai, 206) from Cote d’Ivoire (Coleoptera, Trogidae). Kogane no. 10: 72. Kawai, S. 2010. How to collect Trogids beetle (Coleoptera, Trogidae). Saikaku-Tsûshin no. 20: 31-36 (English title, text in Japanese). Keith, D. 2011. Sur Trox perrisi Fairmaire, 1868 en Turquie (Coleoptera, Scarabaeoidea, Trogidae). Entomologiste (Paris) 67(4): 236. Keith, D., and S. Kawai. 2007. First record of Afromorgus zumpti (Haaf, 1957) from Kenya and Zimbabwe (Coleoptera, Trogidae). Kogane no. 8: 97. Klug, J. F. C. 1868. Coleoptera (pars). In: W. C. H. Peters, Naturwissenschaftliche Reise nach Mossambique auf Befehl seiner Majestät des Königs Friedrich Wilhelm IV in den Jahren 1842 bis 1848 ausgeführt. Reimer; Berlin. 116 p. Koenig, E. 1901. Erster Beitrag zur Coleopteren-Fauna d. Kaukasus. Wiener Entomologische Zeitung 29: 9-10. Kolbe, H. J. 1897. Deutsche Ost-Afrika. Die Thierwelt Ost-Afrikas und Nachburgebiete. Käfer, 4. Reimer; Berlin. 367 p. Kolbe, H. 1910. 18. Scarabaeidae. p. 341-362. In: Y. Sjöstedt (ed.).Wissenschaftliche Ergebnisse der Schwedischen Zoologischen Expedition nach dem Kilimandjaro, dem Meru und den Umbegenden Massaisteppen Deutsch-Ostafrikas 1905-1906. 1. Band. Abteilung 7. Coleoptera. P. Palmquist Artiebolag; Stockholm. 442 p., 4 taf. Kovalenko, Y. N. 2012. New data on the distribution of Trox perrisii (Coleoptera, Trogidae) in Ukraine and bordering Russian territory. Vestnik Zoologii 46(2): 184. Král, D. 1993. Trogidae (p. 67). In: J. Jelínek, Check-list of Czechoslovak insects IV (Coleoptera). Folia Heyrovskyana, Supplementum 1: 1-172. CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 35 Künckel d’Herculais, J. 1887. In: A. Grandidier. Histoire physique, naturelle et politique de Madagascar. Vol. XXII. Histoire naturelle des Coléoptères, Tome II, Atlas, 1ère partie. Imprimerie Nationale; Paris. 25 pls. Laboulbène, A. 1894a. [Note on T. arenarius]. Bulletin des Séances de la Société Entomologique de France 6: 84. Laboulbène, A. 1894b. Communique les observations suivantes: Trox arenarius. Bulletin des Séances de la Société Entomologique de France 6: 134. Lago, P. K., R. L. Post, and C. Y. Oseto. 1979. The phytophagous Scarabaeidae and Troginae of North Dakota. North Dakota Insects, Publication no. 12: 1-131. Latreille, P. A. 1804. Histoire naturelle, génerale et particulière, des Crustacés et des Insectes. Ouvrage faisant suite aux oeuvres de Leclerc de Buffon, et partie du cours complet d’Histoire naturelle rédigé par C. S. Sonnini, membre de plusieurs sociétés savantes. Tome dixièeme. F. Dufart; Paris. 445 p., pls. 81-89. Latreille, P. A. 1825. Familles naturelles du règne animal, exposées succinctement et dans un ordre analytique aves l’indication de leurs genres. Encyclopédie Méthodique, Vol. 10: 345-832. J.- B. Baillière; Paris. LeConte, J. L., and G. H. Horn. 1883. Classification of the Coleoptera of North America. Smithsonian Miscellaneous Collections 26(4): 1-567. Leefmans, S. 1932. Biologische gegevens van een in grotten levenden Trox uit Zuid-Celebes (Trox costatus Wied.). Tijdschrift voor Entomologie (Amsterdam) 75: 36-43. Leng, C. W. 1920. Catalogue of the Coleoptera of America, north of Mexico. J. D. Sherman; Mt. Vernon, New York. 470 p. Leonard, M. D. 1928. A list of the insects of New York. Cornell University Memoir 101. Linné, C. 1761. Fauna suecica sistens Animalia Sueciae Regni: Mammalia, Aves, Amphibia, Pisces, Insecta, Vermes. Distributa per classes et ordines, genera et species. Editio 2. Laurentii Salvii; Stockholm. 578 p. Löbl, I., and A. Smetana. (eds.) 2006. Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea – Scirtoidea – Dascilloidea – Buprestoidea – Byrrhoidea. Apollo Books; Stenstrup. 690 p. López-Colón, J. I. 1991. Nuevos datos sobre la presencia del Trox (Trox) granulipennis Fairmaire, 1852 en la Peninsula Ibérica. Nouvelle Révue d’Entomologie (n. s.) 8: 215. Lumaret, J.-P. 1983. La nidification de Trox. Bulletin de la Société Entomologique de France 88: 594-596. Machatschke, J. W. 1969. Familienreihe Lamellicornia (p. 265-371). In: H. Freude, K. W. Harde, and G. A. Lohse (eds.), Die Käfer Mitteleuropas, Bd. 9. Goecke and Evers; Krefeld. 388 p. Martín-Piera, F., and J. I. López-Colón. 2000. Fauna Iberica Vol. 14, Coleoptera Scarabaeoidea I. Museo Nacional de Ciencias Naturales; Madrid. 526 p. [Trogidae p. 75-93]. Masumoto, K. 1985. Trogidae. In: S.-I. Uéo, Y. Kurosawa, and M. Satô, The Coleoptera of Japan in colour, vol. 2. Hoikusha Publ. Co.; Osaka. 514 p. Medvedev, S. I. 1952. Lichinki plastinchatousykh zhukov. Izdatelstvo Akademii Nauk SSSR; Moskva – Leningrad. 342 p. (in Russian) [Trogidae p. 47-49]. Medvedev, S. I. 1974. Dopolnenie k statie o plastinchatousykh sobrannykh sovietsko-mongolskimi ekspediciami v 1967-1969. Nasekomye Mongolii 2: 110-115. Medvedev, S. I. 1976. Plastinchatousye sobrannye entomologicheskim otriadom sovietsko-mongolskoi biologicheskoi ekspedicii v 1970-71. Nasekomye Mongolii 4: 155-164. Medvedev, S. I., and G. V. Nikolaev. 1972. Descriptions of the immature stages of beetles of the genera Trox F. and Lethrus Scop. and notes on their biology. Entomological Review (Washington, DC), 51: 371-375. [transl.] [Trox larva of Medvedev (1952: 48) most likely is an ochodaeid – see also Nikolaev 1987: 129.] Miksic, R. 1958. Beschreibung neuer und bemerkungen über bekannte Scarabaeiden aus Europa, Asien und Afrika. Bollettino dell’ Associazione Romana di Entomologia 13: 1-18. Mimura, Y. 2001. A new record of Trox mandli from Kyushu, Japan. Kogane no. 2: 65 (in Japanese). Moore, B. P. 1986. A guide to the beetles of southeastern Australia. Australian Entomology Press (Greenwich), Fasc. 7 [Trogidae p. 101-116]. 36 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK Müller, G. 1942. Nuovi coleotteri dell’ Africa orientale. Seconda serie. Atti del Museo Civico di Storia Naturale di Trieste 15(3): 63-86. Müller, O. F. 1776. Zoologiae Danicae prodromus, seu animalium Daniae et Norvegiae indigenarum. Characteres, nomina et synonyma imprimis popularium. Typis Hallageriis; Copenhagen. 282 p. Mulsant, E. 1842. Histoire naturelle des Coléoptères de France. 2. Lamellicornes. Maison Libraire (Paris) & Imprimerie de Dumoulin, Ronet et Sibuet (Lyon); 626 p. Nii, S. 2011. Record of Trox (Coleoptera, Trogidae) from Mt. Gomadanzan, Wakayama prefecture. Saikaku-Tsûshin no. 23: 62 (in Japanese). Nikolaev, G. V. 1977. Zamiatki po sinonimii plastinchatousykh zhukov (Col. Sc.) Mongolii i sopredelnykh teritorii. Nasekomye Mongolii 5: 268-271. Nikolaev, G. V. 1987. Plastinchatousye zhuki (Coleoptera, Scarabaeidae) Kazakhstana i Srednei Azii. Akademia Nauk Kazakhskoi SSR; Alma-Ata. 232 p. Panin, S. 1957. Insecta: Coleoptera, Fam. Scarabaeidae. Fauna Republicii Populare Romine 10(4): 1-315. Panzer, G. W. F. 1795. Entomologia Germanica exhibens insecta per germaniam indigena secundum classes, ordines, genera, species adiectis synonymis, locis, observationibus. I. Eleutherata. Felssecker; Norimbergae. 368 p. Paulian, R. 1943. Les Coléoptères. Payot; Paris. 396 p. Paulian, R. 1944. Hybosoridae et Trogidae. In: Parc National Albert. Mission G. F. de Witte 1933- 1935. Institut des Parcs Nationaux du Congo Belge, Fasc. 46: 1-8. Paulian, R. 1945. Coléoptères Scarabéides de l’Indochine. Première partie. Faune de l’Empire Français, III. Librairie Larose; Paris. 225 p. Paulian, R. 1956. Atlas des larves d’insectes de France. Boubée et Cie; Paris. 222 p. Perris, E. 1877. Larves de Coléoptères. Annales de la Société Linnéenne de Lyon 22: 259-418. Pivotti, I., R. Agoglitta, M. Zunino, E. Piattella, M. Dellacasa, C. Corallini, and D. Mifsud. 2011. The Scarabaeoidea of the Maltese Islands (central Mediterranean) (Coleoptera). Bulletin of the Entomological Society of Malta 4: 85-124. Preyssler, J. D. E. 1790. Verzeichniss böhmischer Insecten. Erstes Hundert. Schoenfeld-Meissner; Prag. 108 p. Ratcliffe, B. C. 1983. Trox hamatus Robinson (Troginae) using a Canthon (Scarabaeinae) brood ball, and new records of North American Trox. Transactions of the Nebraska Academy of Science 11: 53-55. Ratcliffe, B. C. 1991. The scarab beetles of Nebraska. University of Nebraska State Museum, Bulletin 12: 1-333. Ratcliffe, B. C., and M. J. Paulsen. 2008. The scarabaeoid beetles of Nebraska (Coleoptera: Scarabaeoidea). Bulletin of the University of Nebraska State Museum 22: 1-570. Redtenbacher, L. 1858. Fauna Austriaca. Die Käfer. Nach der analytischen Methode bearbeitet. Zweite Auflage. Gerold; Wien. 1017 p. Reitter, E. 1893a. Coleopterologische Notizen. Wiener Entomologische Zeitung 12: 73. Reitter, E. 1893b. Bestimmungs-Tabellen der Lucaniden und coprophagen Lamellicornen der palaearktischen Faunengebietes. Verhandlungen des Naturforschenden Vereins in Brünn 31[1892]: 1-109. Reymond, M. A. 1933. Note sur les deux Scarabaeidae characteristiques récoltés en Mongolie intérieure ou cours de la mission Citroën centre Asie. Bulletin du Muséum National d’Histoire Naturelle (Paris) 5(2): 209-211. Roffey, J. 1958. Observations in the biology of Trox procerus Har., a predator on eggs of the desert locust Schistocerca gregaria (Forsk.). Bulletin of Entomological Research 49: 449-465. Romero-Samper, J., and F. Martín-Piera. 1990. Comportamiento reproductor de Trox perlatus hispanicus Harold, 1872 y Trox hispidus (Pontoppidan, 1763). Elytron 4: 101-109. Schiödte, J. C. 1874. De metamorphosi eleutheratorum observationes. Naturhistorisk Tidsskrift 9: 227-376. Schultze, W. 1916. Beitrag zur coleopteren Fauna der Philippinen, 2. Philippine Journal of Science (D) 11: 291-299. CHECKLIST AND BIBLIOGRAPHY OF THE TROGIDAE INSECTA MUNDI 0314, July 2013 • 37 Scriba, L. G. 1790a. Verzeichniss der Insekten der Darmstädter Gegend. Journal für die Liebhaber der Entomologie 1: 41-73. Scriba, L. G. 1790b. Beiträge zu der Insekten-Geschichte.Varrentrap und Wenner; Frankfurt a. M. 1: 30-47. Scriba, L. G. 1791. Entomologische Bemerkungen und Erfahrungen. Journal für die Liebhaber der Entomologie 3: 244-255. Sharp, D. 1897. On the stridulatory organs of Trox. The Entomologist’s Monthly Magazine 33: 206-207. Smith, A. B. T. 2006. A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a current classification (p. 144-204). In: M. L. Jameson, and B. C. Ratcliffe. (eds.), Scarabaeoidea in the 21st Century: A Festschrift Honoring Henry F. Howden. Coleopterists Society Monograph no. 5: 216 p. Smith, S. G., and N. Virkki. 1978. Animal cytogenetics, vol. 3: Insecta 5. Coleoptera. Gebrüder Bontraeger; Berlin und Stuttgart. p. 235-261. Solier, A. J. J. 1851. Coleopteros. p. 1-285. In: C. Gay (ed.), Historia fisica y politica de Chile segun documentos adquiridos en esta Republica durante doce años de residencia en ella y publicada bajo los auspicios del supremo gobierno. Zoologia. Tomo quinta. Paris – Santiago. 563 p. Solsky, S. 1871. Coléoptères de la Sibérie orientale. Horae Societatis Entomologicae Rossicae 7: 334- 406. Spector, W. 1943. Collecting beetles (Trox) with feather bait traps. Entomological News 54: 224- 229. Stebnicka, Z. 1983. Lucanidae - Trogidae. In: Part XIX (26-27): Coleoptera. Keys for identification of Polish insects. Polish Scientific Publishers, Warsaw. 26 p. Stephens, J. F. 1830. Illustrations of British entomology, or a synopsis of indigenous insects, containing their generic and specific distinctions, with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata. Vol. 3. Baldwin and Cradock; London. 374 p. Strand, A. 1959. Coleoptera i reir av musvak (Buteo buteo L.). Norsk Entomologisk Tidsskrift 11: 46-49. Sturm, J. 1807. Deutschlands Fauna in Abbildungen nach der Natur mit Beschreibungen. V. Abtheilung. Die Insecten. Zweites Bändchen. Käfer. Verfasser; Nürnberg. 279 p. Tesar, Z. 1957. Fauna CSR, 11: Brouci listorozí. Lamellicornia, 2. Scarabaeidae Laparosticti. Ceskoslovenská Akademie Ved; Praha. 336 p. Thomson, C. G. 1863. Skandinaviens Coleoptera, synoptisk bearbetade. Tom V. Lundebergska Boktryckeriet; Lund. 340 p. Thunberg, C. P. 1818. Coleoptera Capensia, antennis lamellatis, sive clava fissili instructa. Mémoires de l’Académie Impériale des Sciences de St. Pétersbourg 6[1813-1814]: 395-450. Van Dyke, E. C. 1928. Notes and descriptions of new species of Scarabaeidae from western North America. Pan-Pacific Entomologist 4: 151-162. Van Emden, F. I. 1941. Larvae of British beetles, 2. A key to the British Lamellicornia larvae. The Entomologist’s Monthly Magazine 77: 117-127, 181-192. Van Emden, F. L. 1948. A Trox larva feeding on locust eggs in Somalia. Proceedings of the Royal Entomological Society of London (B) 17: 145-148. Vaurie, P. 1958. New distribution records of North American Trox. The Coleopterists Bulletin 12: 43-46. Virkki, N. 1967. Chromosome relationships in some North American scarabaeid beetles, with special reference to Pleocoma and Trox. Canadian Journal of Genetics and Cytology 9: 107-125. Voet, J. E. 1776. Catalogus systematicus Coleopterorum. Catalogue systématique des Coléoptères. Systematische naamlyst van dat geslacht van Insecten dat men Torren noemt. Tomus I. Bakhuysen; Den Haag. p. 1-74 (Latin), 1-114 (French), 1-111 (Dutch), 55 pls. Waterhouse, G. R. 1834. Descriptions of the larvae and pupae of various species of coleopterous insects. Transactions of the Royal Entomological Society of London 1: 27-34. Westwood, J. O. 1846. On the lamellicorn beetles which possess extended mandibles and labrum, and 10-jointed antennae. Transactions of the Royal Entomological Society of London 4: 161-180. Xambeu, V. 1892. Moeurs et metamorphose d’insectes. Annales de la Société Linnéenne de Lyon 39: 135-185. 38 • INSECTA MUNDI 0314, July 2013 JIRI ZÍDEK Xambeu, V. 1895. Moeurs et metamorphose d’insectes. Annales de la Société Linnéenne de Lyon 41: 33-84. Xambeu, V. 1896. Moeurs et metamorphose d’insectes; cinquième mémoire (suite). Annales de la Société Linnéenne de Lyon 42: 53-100, 123-188. Yadav, J. S., and R. K. Pillai. 1976. Karyotype orthoselection in Trox. Chromosome Info-Service 20: 8-9. Yadav, J. S., and R. K. Pillai. 1978. Cytotaxonomy of Trox Fabr. Current Science 47(11): 393. Zoubkoff [= Zubkov], V. P. 1833. Nouveaux Coléoptères recueillis Turcménie. Bulletin de la Société Impériale des Naturalistes de Moscou 6: 310-340. Received April 1, 2013; Accepted May 27, 2013.